Physaria chambersii |
Physaria sessilis |
|
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
sessile bladderpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Annuals or biennials; with a fine taproot; densely pubescent, trichomes (sessile or short-stalked), 4–6-rayed, rays usually furcate, rarely bifurcate, (moderately tuberculate or nearly smooth). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
simple or few to several from base, erect, (often distal 1/2 branched), to 6 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade oblanceolate, to 9 cm, margins entire, dentate, or sinuate to lyrate-pinnatifid. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(proximal shortly petiolate, distal sessile); blade narrowly elliptic to linear, 2–4(–6) cm, margins entire or repand to shallowly dentate. |
Racemes | congested. |
loose. |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals elliptic or elongate-ovate, 3.4–5.2(–6.5) mm, (lateral pair subsaccate, median pair thickened apically, cucullate); petals obovate or deltate, 5–10 mm, (sometimes with short claw, margins undulate). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(divaricate-ascending to widely spreading, straight), 8–20 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
globose or subglobose, sometimes slightly compressed, 3–6 mm; valves (not retaining seeds after dehiscence), densely papillose, densely pubescent inside, trichomes raised; replum as wide as or wider than fruit; ovules 8–18 per ovary; style 1.5–3.5 mm. |
Seeds | flattened. |
flattened. |
2n | = 8, 10, 16, 24. |
= 12. |
Physaria chambersii |
Physaria sessilis |
|
Phenology | Flowering Apr–Jul. | Flowering and fruiting Apr–Jun. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Limestone chip, black soils, grassy roadsides, fields, limestone, oak woodlands, mesquite brush lands, pastures, open dry hills |
Elevation | 1500-3200 m (4900-10500 ft) | 30-700 m (100-2300 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
|
TX |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 663. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella gracilis var. sessilis, Lesquerella sessilis |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 328. (2002) |
Web links |