Physaria chambersii |
Physaria scrotiformis |
|
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
silver twinpod, west silver bladderpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; (diminutive); caudex simple or branched, (buried, with thatch of persistent leaf bases distally); (appearing silvery gray-green to silvery purple), densely pubescent, trichomes usually 5 or 6 (rarely 7)-rayed, rays bifurcate or incompletely so, (relatively short, stout, umbonate, moderately tuberculate to nearly smooth, lower layer smoother). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
1–5 from base, prostrate to slightly decumbent, (arising laterally, also erect or ascending from tuft of basal leaves, unbranched, purple-green), 0.08–0.3 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
(petiole slightly winged); blade oblanceolate, elliptic, or rhombic, (mostly flat, sometimes somewhat folded), 0.6–2.7 cm, (base tapering to petiole), margins entire, (apex rounded to rounded-acute). |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(3–7, shortly petiolate or sessile); blade elliptic to oblanceolate, 0.3–0.5 cm, margins entire. |
Racemes | congested. |
crowded, (ca. 3–7 fruits). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (greenish yellow), linear-triangular, 3.7–5 mm, (lateral pair subsaccate); petals oblanceolate to narrowly obovate, 4.5–9 mm. |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(ascending, straight), 1.8–3.4 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(shortly stipitate, purple or greenish purple in age), slightly didymous, ovoid to obpyriform, 3–5 mm (wider than long, base rounded-obtuse, apex rounded, flattened, or slightly emarginate); valves (inflated, slightly wider than replum), pubescent, trichomes scattered; replum obovate to orbicular-obdeltate, apex rounded, obtuse, or truncate; septum complete or medially small-perforate; ovules 4–6(–8) per ovary; style 2–3.6 mm. |
Seeds | flattened. |
relatively plump, (ovate to suborbicular, usually rounded on one side, ± flat or concave on the other, not mucilaginous when wetted). |
2n | = 8, 10, 16, 24. |
|
Physaria chambersii |
Physaria scrotiformis |
|
Phenology | Flowering Apr–Jul. | Flowering Jun-early Jul. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Tundra areas with islands of Engelmann spruce on Leadville limestone, amidst limestone cobbles and gravel |
Elevation | 1500-3200 m (4900-10500 ft) | 3500-3700 m (11500-12100 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
|
CO |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria scrotiformis is known only from La Plata County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 662. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | O’Kane: Novon 17: 376, fig. 1. (2007) |
Web links |