Physaria chambersii |
Physaria saximontana |
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Fremont County twinpod, Rocky Mountain or Fremont County twinpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex usually simple; (silvery) pubescent throughout, trichome rays furcate. | ||||
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
several from base, prostrate to decumbent, 0.3–1 dm. |
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Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
(rosulate; petiole winged); blade orbicular to broadly obovate, 1.5–3 cm, margins entire or with broad, obscure toothlike angles each side at apex, (apex obtuse, surfaces densely pubescent, trichomes appressed). |
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Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
blade broadly spatulate to linear-oblanceolate, 1–1.5 cm, margins entire. |
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Racemes | congested. |
condensed, (subumbellate to slightly more elongated, few-flowered). |
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Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (yellowish, often with some purple), narrowly lanceolate, 5–6 mm; petals spatulate, 7.3–9.2 mm, (not clawed). |
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Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(divaricate-ascending, straight to slightly curved), 6–10 mm. |
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Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
didymous, irregular, suborbicular, deeply bilobed, inflated in age, 10–12 × 12–15 mm, (papery, basal sinus absent or obsolete, apical sinus deep); valves (retaining seeds after dehiscence), densely pubescent, trichomes spreading, (ovaries and immature fruit downy); replum narrowly ovate to broadly oblong, not narrowed at middle, as wide as or wider than fruit, apex acute to obtuse; ovules 4 per ovary; style 3–7 mm. |
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Seeds | flattened. |
flattened. |
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2n | = 8, 10, 16, 24. |
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Physaria chambersii |
Physaria saximontana |
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Phenology | Flowering Apr–Jul. | |||||
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | |||||
Elevation | 1500-3200 m (4900-10500 ft) | |||||
Distribution |
AZ; CA; ID; NV; OR; UT
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MT; WY |
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). Physaria saximontana (especially subsp. dentata) is morphologically similar to 22. P. didymocarpa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 661. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | |||||
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | Rollins: Contr. Gray Herb. 214: 13. (1984) | ||||
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