Physaria chambersii |
Physaria rollinsii |
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Rollins' twinpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; (compact); caudex usually simple, (cespitose); (silvery) pubescent throughout, trichomes 6–8-rayed, rays furcate near base, fused at base, (umbonate, strongly tuberculate throughout). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
several from base, decumbent, (unbranched, slender), 0.5–1 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
(strongly rosulate); blade usually oblanceolate or broader, sometimes triangular, 2–3.5 cm (width 5–10 mm), margins entire or with 1 or 2 broad teeth, (apex acute). |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
blade oblanceolate, 1–1.5 cm (width 2–4 mm), margins entire, (apex acute). |
Racemes | congested. |
congested, (elongated moderately in fruit). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals linear, 5–7 mm; petals spatulate, 8–10 mm, (apex often somewhat truncate). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(spreading, straight or somewhat sigmoid), 5–8 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(erect), didymous, suborbicular, inflated, 2–5(–8) × 4–8(–10)mm, (coriaceous, base slightly cordate or nearly obtuse, sinus obsolete or absent, apical sinus broad and deep); valves (retaining seeds after dehiscence), pubescent, trichomes appressed, (silvery on ovaries and immature fruit); replum obovate to oblong, as wide as or wider than fruit, rarely somewhat constricted basally, apex obtuse; ovules 4 per ovary; style 5–7 mm. |
Seeds | flattened. |
slightly flattened. |
2n | = 8, 10, 16, 24. |
= 8. |
Physaria chambersii |
Physaria rollinsii |
|
Phenology | Flowering Apr–Jul. | Flowering May–Jun. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Sagebrush, granitic talus, open knolls, limestone chiprock, steep slopes, clay banks, near granite boulders |
Elevation | 1500-3200 m (4900-10500 ft) | 2300-2500(-3900) m (7500-8200(-12800) ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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CO
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 661. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | G. A. Mulligan: Canad. J. Bot. 44: 1663, fig. 2, plate 1, fig. 4. (1966) |
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