Physaria chambersii |
Physaria reediana |
|
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
alpine bladderpod, reed's twinpod, Rollins' bladderpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex simple or branched, (covered with persistent leaf bases, loosely cespitose); densely pubescent, trichomes (appressed to ascending, plant appearing shaggy, always appressed on fruits), 4- or 5-rayed, rays furcate or bifurcate, (tuberculate throughout). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
few to several from base, ± erect, (arising laterally, also from within basal leaves), 0.2–0.4 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
(erect); blade linear-oblanceolate, 1.2–2.8 cm, (base gradually narrowed to petiole), margins entire. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
blade linear, similar to basal. |
Racemes | congested. |
dense, (often subumbellate, not or barely exceeding basal leaves). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (pale green-yellow), oblong to elliptic, 4–5 mm, (median pair usually thickened apically, cucullate); petals lingulate, 6–9 mm. |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(ascending, curved), 3–5.5 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
lanceolate in outline, compressed (latiseptate) on margins and at apex, 4–5 mm; valves pubescent, trichomes closely appressed; ovules 8–12 per ovary; style 3.5–4.5 mm (equaling or exceeding length of fruit, curved proximal to stigma). |
Seeds | flattened. |
plump, (oblong). |
2n | = 8, 10, 16, 24. |
|
Physaria chambersii |
Physaria reediana |
|
Phenology | Flowering Apr–Jul. | Flowering May–Jul. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Open areas of grasslands on calcareous soils |
Elevation | 1500-3200 m (4900-10500 ft) | 1200-1900 m (3900-6200 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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CO; NE; WY
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 660. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Vesicaria alpina, Alyssum alpinum, Lesquerella alpina, Lesquerella alpina var. laevis, Lesquerella condensata var. laevis |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | O’Kane & Al-Shehbaz: Novon 12: 327. (2002) |
Web links |