Physaria chambersii |
Physaria pycnantha |
|
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
mountain-view bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex branched, (densely cespitose and forming hemispheric mounds); densely pubescent, trichomes 5-rayed, rays bifurcate near base, fused at base, (strongly tuberculate throughout). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
few to several from base, erect, (usually exceeding basal leaves), 0.3–0.7 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade linear-spatulate, 1.5–4 cm, (base narrowed gradually to petiole), margins entire. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
blade spatulate, similar to basal. |
Racemes | congested. |
crowded in distal 1/3, (4–10-flowered). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (pale yellow), oblong to elliptic, 3–4 mm, (median pair usually thickened apically, cucullate); petals (sometimes with slight tinge of orange basally), lingulate, 4–6 mm. |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(loosely to strongly sigmoid), 6–10 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
ellipsoid, slightly inflated (somewhat latiseptate), 4–5 mm, (apex acute); valves pubescent, trichomes erect, appearing slightly shaggy; ovules 4–8 per ovary; styles 2.5–3 mm, (shorter than mature fruits). |
Seeds | flattened. |
± flattened, convex on outer side. |
2n | = 8, 10, 16, 24. |
|
Physaria chambersii |
Physaria pycnantha |
|
Phenology | Flowering Apr–Jul. | Flowering late May–Jun(-Jul). |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Dry, windswept knolls of limestone gravel, with other cushion-forming plants |
Elevation | 1500-3200 m (4900-10500 ft) | 1600-2300 m (5200-7500 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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ID; MT
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria pycnantha is morphologically similar to 56. P. nelsonii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 659. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | Grady & O’Kane: Novon 17: 188, fig. 5. (2007) |
Web links |