Physaria chambersii |
Physaria purpurea |
|
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
rose bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex simple, (usually woody); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, (smooth or tuberculate). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
simple from base, erect, (unbranched, sparsely leaved), to 7 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade elliptic or obovate to oblong, 4–15 cm, margins entire, dentate, or lyrate-pinnatifid. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(proximal often narrowed to short petiole, distal sessile); blade broadly elliptic to obovate or rhombic, 0.5–3(–5) cm, margins entire. |
Racemes | congested. |
dense or slightly elongated. |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals elliptic to ovate, 3.5–6(–7) mm, (median pair usually thickened apically, cucullate); petals (white, often purple-veined, fading purplish), suborbicular to obovate, obdeltate, or cuneate, 4.5–10(–12) mm, (often narrowed to broad claw, apex emarginate, less frequently claw undifferentiated from blade). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(spreading or recurved, loosely sigmoid), 5–25 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(pendent or horizontal, sessile or substipitate), subglobose to broadly ellipsoid, not or slightly inflated, (4–)5–8 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–8(–12) per ovary; style 1–3(–4) mm. |
Seeds | flattened. |
flattened. |
2n | = 8, 10, 16, 24. |
= 18, 36. |
Physaria chambersii |
Physaria purpurea |
|
Phenology | Flowering Apr–Jul. | Flowering Mar–Oct. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Rocky draws, canyons, stony hills, ridges, rock crevices on limestone ledges, lava cliffs, sand and gravel of dry stream beds, rocky slopes, talus, shade of bushes or cactus clumps |
Elevation | 1500-3200 m (4900-10500 ft) | 400-2400 m (1300-7900 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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AZ; NM; TX; Mexico (Chihuahua, Coahuila, Sonora)
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 659. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Vesicaria purpurea, Lesquerella purpurea, Lesquerella purpurea subsp. foliosa, Lesquerella purpurea var. foliosa, P. purpurea var. foliosa |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 327. (2002) |
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