Physaria chambersii |
Physaria pulvinata |
|
---|---|---|
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
cushion bladderpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex (buried), branched, (dense, forming hard mats); densely pubescent, trichomes (subsessile), 8–13-rayed, rays usually furcate, distinct, (umbonate, usually tuberculate, less so over umbo). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
several (to several hundred) from base, erect, (each terminating in a tufted cluster of leaves), to 7 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
(petiole not differentiated from blade); blade narrowly elliptic to narrowly linear-oblanceolate, (0.8–)1–1.5 cm, (base cuneate), margins entire. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
similar to basal, blade sometimes linear, (apex acute). |
Racemes | congested. |
dense, (often ± subumbellate, somewhat elongated in fruit). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals narrowly elliptic, 2.5–3.5(–4) mm, (not keeled); petals narrowly spatulate, 4–7 mm. |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(strongly sigmoid), 5–10 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
ellipsoid, compressed, 4–6 mm; valves densely pubescent, trichomes appressed; ovules 2 per ovary; style 2–3.5 mm. |
Seeds | flattened. |
flattened, (oval). |
2n | = 8, 10, 16, 24. |
|
Physaria chambersii |
Physaria pulvinata |
|
Phenology | Flowering Apr–Jul. | Flowering late May–Jun. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Gray, argillaceous shale outcrops with sagebrush and junipers |
Elevation | 1500-3200 m (4900-10500 ft) | 2300-2600 m (7500-8500 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
|
CO |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria pulvinata is known from an area surrounded by a pygmy forest of Utah juniper in Dolores and San Miguel Counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 659. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | O’Kane & Reveal: Brittonia 58: 74, fig. 1. (2006) |
Web links |