Physaria chambersii |
Physaria prostrata |
|
---|---|---|
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
low bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex branched; densely pubescent, trichomes (usually sessile, rarely short-stalked), several-rayed, rays furcate or bifurcate, (umbonate, tuberculate throughout). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
several from base, usually prostrate, rarely decumbent, (unbranched, often purplish, sparsely pubescent), to 1.5 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade deltate, hastate, or, less often, rhombic to elliptic, 1–5 cm, margins entire (often partially involute). |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(proximal shortly petiolate); blade linear to oblanceolate, 0.5–1.5 cm, margins entire. |
Racemes | congested. |
loose, elongated. |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (often purplish), oblong, lanceolate, or ovate, 4–6 mm, (median pair thickened apically); petals spatulate or cuneate, 5–8(–9) mm, (margins undulate). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(ascending, slightly sigmoid to straight), 5–10 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
ovoid or ellipsoid, slightly compressed, 3–5(–6) mm, (base often gibbous); valves pubescent, trichomes loose, furcate near their bases and spreading, sometimes sparsely pubescent inside; ovules 4(–8) per ovary; style 1.5–4 mm. |
Seeds | flattened. |
flattened. |
2n | = 8, 10, 16, 24. |
|
Physaria chambersii |
Physaria prostrata |
|
Phenology | Flowering Apr–Jul. | Flowering May–Jun. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Whitish sand and small rocks on steep slopes, dry hillsides, windswept knolls, shaley slopes |
Elevation | 1500-3200 m (4900-10500 ft) | 1800-2500 m (5900-8200 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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ID; UT; WY |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria prostrata is sometimes found on igneous substrates, which is unusual for the genus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 658. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella prostrata |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (A. Nelson) O’Kane & Al-Shehbaz: Novon 12: 327. (2002) |
Web links |