Physaria chambersii |
Physaria pinetorum |
|
---|---|---|
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
White Mountain bladderpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 6–8-rayed, rays furcate or bifurcate, (tuberculate, less so on outer layers). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
simple or few from base, ascending to erect, (0.5–)1–2(–3.5) dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
(petiole tapering to blade); blade rhombic to elliptic and irregularly angular, sometimes spatulate to oblanceolate, 1.5–7.5(–10) cm, margins entire. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(not or loosely overlapping, petiolate or distal sessile); blade spatulate to oblanceolate, 1–4 cm, margins entire. |
Racemes | congested. |
crowded, elongated. |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals ovate, oblong, or elliptic 4–7.5 mm, (median pair thickened apically, cucullate); petals spatulate or broadly cuneate, 6–13 mm, (claw slightly expanded at base). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(ascending, curved or sigmoid), 6–12(–20) mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(substipitate), globose or obovoid to ellipsoid, sometimes slightly obcompressed, 4–9 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–24 per ovary; style (2–)4–7 mm. |
Seeds | flattened. |
flattened. |
2n | = 8, 10, 16, 24. |
= 10. |
Physaria chambersii |
Physaria pinetorum |
|
Phenology | Flowering Apr–Jul. | Flowering Apr–Jul. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Scrub oak, pinyon-juniper woodland, open ponderosa pine forests, these sometimes mixed with Douglas fir, white pine, white fir, Engelmann spruce, or Gambel oak, on limestone-derived or otherwise basic soils, often in rock crevices |
Elevation | 1500-3200 m (4900-10500 ft) | 1400-2900 (-3400) m (4600-9500 (-11200) ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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AZ; NM
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria pinetorum with reduced forms are found at high elevations; in disturbed, moist soils plants can become quite large, as in the Manzano Mountains. Densely cespitose plants with crowded racemes not exceeding the basal leaves are found at the crest (3200–3400 m) of the Sandia Mountains, New Mexico. These probably represent an undescribed taxon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 658. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella pinetorum |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (Wooton & Standley) O’Kane & Al-Shehbaz: Novon 12: 327. (2002) |
Web links |