The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod

White Mountain bladderpod

Habit Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 6–8-rayed, rays furcate or bifurcate, (tuberculate, less so on outer layers).
Stems

several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm.

simple or few from base, ascending to erect, (0.5–)1–2(–3.5) dm.

Basal leaves

(petiole slender);

blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate.

(petiole tapering to blade);

blade rhombic to elliptic and irregularly angular, sometimes spatulate to oblanceolate, 1.5–7.5(–10) cm, margins entire.

Cauline leaves

blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute).

(not or loosely overlapping, petiolate or distal sessile);

blade spatulate to oblanceolate, 1–4 cm, margins entire.

Racemes

congested.

crowded, elongated.

Flowers

sepals narrowly lanceolate, 5–8(–9) mm;

petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade).

sepals ovate, oblong, or elliptic 4–7.5 mm, (median pair thickened apically, cucullate);

petals spatulate or broadly cuneate, 6–13 mm, (claw slightly expanded at base).

Fruiting pedicels

(divaricate, slightly sigmoid), 8–15 mm.

(ascending, curved or sigmoid), 6–12(–20) mm.

Fruits

(often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded);

valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent;

replum oblong, as wide as or wider than fruits, apex obtuse;

ovules 4–12 per ovary;

style (4–)6–8 mm (exceeding sinus).

(substipitate), globose or obovoid to ellipsoid, sometimes slightly obcompressed, 4–9 mm;

valves (not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules 4–24 per ovary;

style (2–)4–7 mm.

Seeds

flattened.

flattened.

2n

= 8, 10, 16, 24.

= 10.

Physaria chambersii

Physaria pinetorum

Phenology Flowering Apr–Jul. Flowering Apr–Jul.
Habitat Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas Scrub oak, pinyon-juniper woodland, open ponderosa pine forests, these sometimes mixed with Douglas fir, white pine, white fir, Engelmann spruce, or Gambel oak, on limestone-derived or otherwise basic soils, often in rock crevices
Elevation 1500-3200 m (4900-10500 ft) 1400-2900 (-3400) m (4600-9500 (-11200) ft)
Distribution
from FNA
AZ; CA; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM
[WildflowerSearch map]
[BONAP county map]
Discussion

Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Physaria pinetorum with reduced forms are found at high elevations; in disturbed, moist soils plants can become quite large, as in the Manzano Mountains. Densely cespitose plants with crowded racemes not exceeding the basal leaves are found at the crest (3200–3400 m) of the Sandia Mountains, New Mexico. These probably represent an undescribed taxon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 631. FNA vol. 7, p. 658.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms P. chambersii var. canaani, P. chambersii var. sobolifera Lesquerella pinetorum
Name authority Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) (Wooton & Standley) O’Kane & Al-Shehbaz: Novon 12: 327. (2002)
Web links