Physaria chambersii |
Physaria parvula |
|
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
pygmy bladderpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex (buried), usually branched, sometimes simple, (cespitose); densely pubescent, trichomes (appressed), 4–7-rayed, rays distinct, furcate or bifurcate near base. |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
few to several from base, erect, (unbranched, slender), 0.3–1.5(–3) dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
(tufted, erect); blade linear to very narrowly spatulate, 1–3(–4) cm, margins entire (involute). |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
similar to basal. |
Racemes | congested. |
relatively dense. |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (greenish yellow), elliptic, 3.5–7 mm; petals spatulate, 5–6 mm, (not clawed). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(ascending, curved or sigmoid), 2–10 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(erect), ovoid (or longer than broad), usually inflated, 4–5 mm, (apex acute, slightly flattened); valves pubescent, trichomes appressed; ovules 4–8 per ovary; style 2–4 mm. |
Seeds | flattened. |
flattened, (mucilaginous). |
2n | = 8, 10, 16, 24. |
= 10, 20. |
Physaria chambersii |
Physaria parvula |
|
Phenology | Flowering Apr–Jul. | Flowering May–Jul. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Exposed windblown ridges, gravelly hills, open rocky knolls, gravelly hilltops, clay hillsides, granitic sand, reddish soil, sagebrush, mountain scrub, and pinyon-juniper areas |
Elevation | 1500-3200 m (4900-10500 ft) | 1800-2800 m (5900-9200 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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CO; UT; WY
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 657. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella parvula, Lesquerella alpina subsp. parvula, Lesquerella alpina var. parvula |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (Greene) O’Kane & Al-Shehbaz: Novon 12: 326. (2002) |
Web links |