Physaria chambersii |
Physaria pachyphylla |
|
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
thick-leaf bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex usually simple, rarely branched, (loosely mounded, rosette-like growth); densely (silvery or gray) pubescent, trichomes (sessile), 5-rayed, rays bifurcate, slightly fused near base of main rays, (tuberculate throughout, less over umbo). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
several from base, decumbent to prostrate, (well-exserted beyond basal leaves), 0.2–0.5 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
(petiole differentiated from blade); blade (slightly cupped, leathery, nearly 1 mm thick), oblanceolate to orbicular, 1.2–2 cm, margins entire, (apex acute). |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
blade spatulate, similar to basal. |
Racemes | congested. |
dense, (subumbellate). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (pale yellow), elliptic to oblong, 3.5–4.0 mm, (median pair somewhat thickened apically, cucullate); petals lingulate, 5–6 mm. |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(ascending, curved), 5–7 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
globose or ellipsoid, slightly inflated (with slight apical constriction), 3–6 mm; valves pubescent, trichomes closely appressed; ovules 8 per ovary; style 1–3 mm (shorter than mature fruit). |
Seeds | flattened. |
plump, (oblong). |
2n | = 8, 10, 16, 24. |
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Physaria chambersii |
Physaria pachyphylla |
|
Phenology | Flowering Apr–Jul. | Flowering Jun–Jul. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Barren areas of mixed white, pink, or reddish limestone and diatomaceous earth |
Elevation | 1500-3200 m (4900-10500 ft) | 1300-1600 m (4300-5200 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
|
MT |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria pachyphylla is known from the Pryor Mountain Desert near the Wyoming state line. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 656. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | O’Kane & Grady: Novon 17: 187, fig. 4. (2007) |
Web links |