Physaria chambersii |
Physaria ovalifolia |
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
roundleaf bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex simple or branched, (thickened by persistent leaf bases); densely pubescent (foliage usually scabrous), trichomes (sessile or short-stalked), several-rayed, rays furcate near base, (usually strongly umbonate, roughly tuberculate, less so over umbo). | ||||
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
few to several from base, erect or outer decumbent, 0.5–2.5 dm. |
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Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade suborbicular to elliptic or ovate or deltate, 0.5–2(–6.5) cm, margins entire or shallowly dentate. |
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Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(proximal shortly petiolate, distal usually sessile); blade narrowly elliptic or obovate, (0.5–)1–2.5(–4) cm, margins entire. |
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Racemes | congested. |
compact, (± subumbellate to densely corymbiform, elongated or not). |
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Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals ± elliptic, 4.5–7(–8.5) mm, (median pair thickened apically); petals (sometimes white), suborbicular to obovate or obdeltate, 6.5–15 mm, (base narrowing to broad claw, apex sometimes emarginated). |
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Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(usually spreading at right angles, sometimes nearly erect, ± straight), 5–15(–20) mm, (stout). |
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Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(sessile or shortly stipitate, less than 1 mm), subglobose to broadly ellipsoid, inflated or slightly compressed (terete or subterete), (4–)5–8(–9) mm; valves (not retaining seeds after dehiscence), glabrous; replum as wide as or wider than fruit; ovules 8–16 per ovary; style 4–8(–9) mm. |
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Seeds | flattened. |
flattened. |
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2n | = 8, 10, 16, 24. |
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Physaria chambersii |
Physaria ovalifolia |
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Phenology | Flowering Apr–Jul. | |||||
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | |||||
Elevation | 1500-3200 m (4900-10500 ft) | |||||
Distribution |
AZ; CA; ID; NV; OR; UT
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CO; KS; NE; NM; OK; TX
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 655. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella ovalifolia, Lesquerella engelmannii subsp. ovalifolia | ||||
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (Rydberg) O’Kane & Al-Shehbaz: Novon 12: 326. (2002) | ||||
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