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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod

roundleaf bladderpod

Habit Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). Perennials; caudex simple or branched, (thickened by persistent leaf bases); densely pubescent (foliage usually scabrous), trichomes (sessile or short-stalked), several-rayed, rays furcate near base, (usually strongly umbonate, roughly tuberculate, less so over umbo).
Stems

several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm.

few to several from base, erect or outer decumbent, 0.5–2.5 dm.

Basal leaves

(petiole slender);

blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate.

blade suborbicular to elliptic or ovate or deltate, 0.5–2(–6.5) cm, margins entire or shallowly dentate.

Cauline leaves

blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute).

(proximal shortly petiolate, distal usually sessile);

blade narrowly elliptic or obovate, (0.5–)1–2.5(–4) cm, margins entire.

Racemes

congested.

compact, (± subumbellate to densely corymbiform, elongated or not).

Flowers

sepals narrowly lanceolate, 5–8(–9) mm;

petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade).

sepals ± elliptic, 4.5–7(–8.5) mm, (median pair thickened apically);

petals (sometimes white), suborbicular to obovate or obdeltate, 6.5–15 mm, (base narrowing to broad claw, apex sometimes emarginated).

Fruiting pedicels

(divaricate, slightly sigmoid), 8–15 mm.

(usually spreading at right angles, sometimes nearly erect, ± straight), 5–15(–20) mm, (stout).

Fruits

(often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded);

valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent;

replum oblong, as wide as or wider than fruits, apex obtuse;

ovules 4–12 per ovary;

style (4–)6–8 mm (exceeding sinus).

(sessile or shortly stipitate, less than 1 mm), subglobose to broadly ellipsoid, inflated or slightly compressed (terete or subterete), (4–)5–8(–9) mm;

valves (not retaining seeds after dehiscence), glabrous;

replum as wide as or wider than fruit;

ovules 8–16 per ovary;

style 4–8(–9) mm.

Seeds

flattened.

flattened.

2n

= 8, 10, 16, 24.

Physaria chambersii

Physaria ovalifolia

Phenology Flowering Apr–Jul.
Habitat Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas
Elevation 1500-3200 m (4900-10500 ft)
Distribution
from FNA
AZ; CA; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; KS; NE; NM; OK; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Caudices branched (well-developed); petals usually yellow, rarely white, 6.5-12(-14) mm, usually 1.5 times or less as long as sepals; racemes usually not elongated (subumbellate).
subsp. ovalifolia
1. Caudices usually simple; petals white, (9-)11-15 mm, often 2 times as long as sepals; racemes usually elongated.
subsp. alba
Source FNA vol. 7, p. 631. FNA vol. 7, p. 655.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. ovalifolia subsp. alba, P. ovalifolia subsp. ovalifolia
Synonyms P. chambersii var. canaani, P. chambersii var. sobolifera Lesquerella ovalifolia, Lesquerella engelmannii subsp. ovalifolia
Name authority Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) (Rydberg) O’Kane & Al-Shehbaz: Novon 12: 326. (2002)
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