Physaria chambersii |
Physaria oregona |
|
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Oregon twin-pod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex simple, (cespitose); (silvery) pubescent throughout, trichomes (stalked), few-rayed, rays furcate or imperfectly so, (tuberculate throughout). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
several from base, erect or somewhat decumbent, (unbranched), 1–3.5 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
(petiole slender, usually incised or with broad teeth along petiole); blade obovate, 4–6 cm, margins entire. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
blade oblanceolate or broader, 1.5–2.5 cm, margins entire or sparsely dentate, (apex acute). |
Racemes | congested. |
somewhat loose, (5–15 cm). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals oblong, 5–7 mm; petals (lemon yellow), spatulate, 9–12 mm. |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(spreading or ascending, curved, fruits not pendent), 10–20 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
didymous, obreniform, moderately inflated, angustispetate, (8–)10–12(–15) × 10–14(–16) mm, (papery, not keeled, basal sinus absent, apical sinus broad and open); valves (retaining seeds after dehiscence, rounded or irregular), loosely pubescent, trichomes spreading; replum broadly lanceolate, as wide as or wider than fruit, apex acute; ovules 8 per ovary; style 1–2 mm. |
Seeds | flattened. |
flattened. |
2n | = 8, 10, 16, 24. |
= 8. |
Physaria chambersii |
Physaria oregona |
|
Phenology | Flowering Apr–Jul. | Flowering Apr–Jun. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Gravelly banks, stream shores, rocky slopes, dry hillsides, serpentine soils |
Elevation | 1500-3200 m (4900-10500 ft) | 900-1900 m (3000-6200 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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ID; OR; WA
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 655. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Coulterina oregona, Lesquerella oregona |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | S. Watson: Proc. Amer. Acad. Arts 17: 363. (1882) |
Web links |