Physaria chambersii |
Physaria occidentalis |
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
western bladder-pod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex simple or branched, (thickened, sometimes subterranean); densely pubescent, trichomes (sessile or short-stalked), 5–7-rayed, raysdistinct, bifurcate, (moderately to prominently tuberculate). | ||||
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
few to several from base, prostrate to decumbent or erect, (usually unbranched), 0.3–1.5(–3) dm. |
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Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
(petiole slender); blade suborbicular to obovate or elliptic, 1–8 cm, (base narrowing gradually or abruptly to petiole), margins sinuate-dentate or entire. |
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Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(proximal shortly petiolate, distal sessile); blade oblanceolate, 0.5–1.5(–2.5) cm, margins entire or remotely dentate. |
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Racemes | congested. |
dense or loose. |
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Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals elliptic or oblong-elliptic, 4.5–7 mm, (median pair cucullate); petals spatulate, 7–9(–14) mm, (claw undifferentiated from blade). |
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Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(sigmoid or curved), 5–10(–15) mm. |
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Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(erect), ellipsoid to obovoid, compressed at apex and sometimes margins (strongly latiseptate), (5–)6–9 mm, (apex acute, often beaked); valves densely pubescent, trichomes appressed or spreading, sometimes sparsely pubescent inside, trichomes 4- or 5-rayed; ovules 4–12 per ovary; style (2–)3–6.5 mm, (often sparsely pubescent). |
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Seeds | flattened. |
inner surface flattened, outer convex. |
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2n | = 8, 10, 16, 24. |
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Physaria chambersii |
Physaria occidentalis |
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Phenology | Flowering Apr–Jul. | |||||
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | |||||
Elevation | 1500-3200 m (4900-10500 ft) | |||||
Distribution |
AZ; CA; ID; NV; OR; UT
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CA; ID; NV; OR; UT; WA
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 654. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Vesicaria occidentalis, Lesquerella occidentalis | ||||
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 326. (2002) | ||||
Web links |