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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod

western bladder-pod

Habit Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). Perennials; caudex simple or branched, (thickened, sometimes subterranean); densely pubescent, trichomes (sessile or short-stalked), 5–7-rayed, raysdistinct, bifurcate, (moderately to prominently tuberculate).
Stems

several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm.

few to several from base, prostrate to decumbent or erect, (usually unbranched), 0.3–1.5(–3) dm.

Basal leaves

(petiole slender);

blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate.

(petiole slender);

blade suborbicular to obovate or elliptic, 1–8 cm, (base narrowing gradually or abruptly to petiole), margins sinuate-dentate or entire.

Cauline leaves

blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute).

(proximal shortly petiolate, distal sessile);

blade oblanceolate, 0.5–1.5(–2.5) cm, margins entire or remotely dentate.

Racemes

congested.

dense or loose.

Flowers

sepals narrowly lanceolate, 5–8(–9) mm;

petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade).

sepals elliptic or oblong-elliptic, 4.5–7 mm, (median pair cucullate);

petals spatulate, 7–9(–14) mm, (claw undifferentiated from blade).

Fruiting pedicels

(divaricate, slightly sigmoid), 8–15 mm.

(sigmoid or curved), 5–10(–15) mm.

Fruits

(often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded);

valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent;

replum oblong, as wide as or wider than fruits, apex obtuse;

ovules 4–12 per ovary;

style (4–)6–8 mm (exceeding sinus).

(erect), ellipsoid to obovoid, compressed at apex and sometimes margins (strongly latiseptate), (5–)6–9 mm, (apex acute, often beaked);

valves densely pubescent, trichomes appressed or spreading, sometimes sparsely pubescent inside, trichomes 4- or 5-rayed;

ovules 4–12 per ovary;

style (2–)3–6.5 mm, (often sparsely pubescent).

Seeds

flattened.

inner surface flattened, outer convex.

2n

= 8, 10, 16, 24.

Physaria chambersii

Physaria occidentalis

Phenology Flowering Apr–Jul.
Habitat Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas
Elevation 1500-3200 m (4900-10500 ft)
Distribution
from FNA
AZ; CA; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; NV; OR; UT; WA
[WildflowerSearch map]
[BONAP county map]
Discussion

Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems erect or decumbent; fruit margins and apices compressed, apices often beaked; valves pubescent inside; ovules 4-8 per ovary; styles frequently curved; n California, c Idaho, nw Nevada, Oregon, Washington.
subsp. occidentalis
1. Stems prostrate to decumbent; fruit, margins not conspicuously compressed, apices somewhat compressed, usually not conspicuously beaked; valves usually glabrous, sometimes sparsely pubescent inside; ovules 8-12 per ovary; styles usually straight; Nevada, w Utah.
subsp. cinerascens
Source FNA vol. 7, p. 631. FNA vol. 7, p. 654.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. occidentalis subsp. cinerascens, P. occidentalis subsp. occidentalis
Synonyms P. chambersii var. canaani, P. chambersii var. sobolifera Vesicaria occidentalis, Lesquerella occidentalis
Name authority Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 326. (2002)
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