Physaria chambersii |
Physaria obcordata |
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Dudley bluffs twinpod, Piceance twinpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex branched, (thick, covered with persistent, overlapping leaf bases); densely (silvery) pubescent, trichomes (stellate-scalelike), several-rayed, rays fused (webbed) to tips. |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
several from base, erect to decumbent, (unbranched, fertile stems from among basal leaves), 1.2–1.8 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
(erect, not rosulate); blade broadly oblanceolate, 4–8 cm, margins entire or shallowly sinuate-dentate, (apex acute). |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(proximal petiolate); blade narrowly lanceolate, similar to basal in size, (distal with cuneate base), margins entire. |
Racemes | congested. |
loose, (elongated in fruit). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (greenish yellow), elliptic, often broadly so, 4.8–7.1 mm; petals oblanceolate, 6.8–9.8 mm. |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(widely spreading to recurved), 1–1.5 cm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(usually pendent), slightly didymous apically, obcordate, slightly inflated, 4–7 × 3–6 mm, (papery, basal sinus absent, apical sinus evident to nearly absent); valves (retaining seeds after dehiscence), pubescent; replum broadly obovate to suborbicular, 4–5 mm, as wide as or wider than fruit, margins entire; ovules usually 4 (rarely 6–8) per ovary; style (2.5–)3–4(–5) mm. |
Seeds | flattened. |
plump. |
2n | = 8, 10, 16, 24. |
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Physaria chambersii |
Physaria obcordata |
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Phenology | Flowering Apr–Jul. | Flowering May–Jun. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Steep slopes, fine chiprock, shaley hillsides |
Elevation | 1500-3200 m (4900-10500 ft) | 1800-2300 m (5900-7500 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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CO
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria obcordata is known from the Thirteenmile Creek Tongue and the Parachute Creek Member of the Green River Formation. It is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 653. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | Rollins: J. Arnold Arbor. 64: 495, fig. 1. (1983) |
Web links |