Physaria chambersii |
Physaria nelsonii |
|
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Nelson's bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; (diminutive); caudex branched, (densely cespitose, mound-forming); densely pubescent, trichomes (flaring, giving a shaggy appearance), usually 4–5-rayed, rays furcate or bifurcate, (strongly tuberculate throughout). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
few to several from base, erect, (from basal tuft), 0.1–0.2(–0.3) dm, (not or just barely exceeding leaves). |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade spatulate to oblanceolate, 0.5–1.5 cm, (base gradually narrowed to petiole), margins entire. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(absent or few); similar to basal, blade linear. |
Racemes | congested. |
dense, (few-flowered). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (pale yellow), oblong to elliptic, 4–5 mm, (median pair usually thickened apically, cucullate); petals (bright yellow), lingulate, 6–8 mm. |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(loosely sigmoid), 3–5 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
lanceolate, compressed apically, 2.5–3(–4) mm; valves pubescent, trichomes spreading, (appearing fuzzy), rarely with trichomes inside; ovules 4–8 per ovary; style 2.5–4 mm. |
Seeds | flattened. |
plump, (oblong). |
2n | = 8, 10, 16, 24. |
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Physaria chambersii |
Physaria nelsonii |
|
Phenology | Flowering Apr–Jul. | Flowering May-mid Jun. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Limestone, windswept knolls and cliffs, nearly barren areas with other cushion-forming plants |
Elevation | 1500-3200 m (4900-10500 ft) | 1600-2300 m (5200-7500 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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UT; WY |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria nelsonii is morphologically similar to 73. P. pycnantha, which traditionally was included in a broader P. nelsonii. These allopatric species are distinguished by styles equal to or exceeding the length of fruit and fruiting stems overtopped by basal leaves (P. nelsonii) versus styles shorter than fruits and fruiting stems usually exserted beyond basal leaves (P. pycnantha). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 652. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella condensata, Lesquerella alpina subsp. condensata, Lesquerella alpina var. condensata |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | O’Kane & Al-Shehbaz: Novon 12: 326. (2002) |
Web links |