Physaria chambersii |
Physaria navajoensis |
|
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Navajo bladderpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex branched, (woody, pulvinate-cespitose, forming hard, hemispherical mats, basal parts covered with persistent leaf bases); densely (silvery gray) pubescent, trichomes mostly 5-rayed, rays bifurcate, slightly fused at base, (umbonate, strongly tuberculate except nearly smooth over umbo). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
several from base (crowded), erect, not exceeding leaves. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
usually absent. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(petiole not differentiated from blade); blade linear-oblanceolate, 3–8(–13) mm, margins entire. |
Racemes | congested. |
(secund), dense, corymbose, (few-flowered, not or barely exceeding leaves). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (yellow-green), linear to narrowly triangular, 3.7–4.8 mm, (lateral pair subsaccate); petals (deep yellow, slightly orange in center), spatulate, 5.2–6.5 mm, (claw joined at right angle). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(ascending to divaricate-ascending, straight), 3.5–6 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(becoming reddish or copper-colored in age), ovate, often slightly compressed (at margins apically), 3–5 mm, (apex acute); valves (not retaining seeds after dehiscence), glabrous; (septum perforate or not); ovules 4–8 per ovary; style 1.8–3 mm. |
Seeds | flattened. |
plump or slightly flattened, (strongly mucilaginous). |
2n | = 8, 10, 16, 24. |
|
Physaria chambersii |
Physaria navajoensis |
|
Phenology | Flowering Apr–Jul. | Flowering May-early Jun. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Pinyon-juniper communities on nearly barren outcrops of Todilto Limestone |
Elevation | 1500-3200 m (4900-10500 ft) | 2200-2400 m (7200-7900 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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AZ; NM |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria navajoensis is morphologically similar to 85. P. tumulosa of southern Utah, differing subtly. Physaria navajoensis has petals slightly orange at the junction of blade and claw, a sharp bend at that junction giving the flower a flat-topped appearance, and strongly mucilaginous seeds. Physaria tumulosa has pure yellow petals that gently flex at the junction of blade and claw, and seeds that are not mucilaginous. Molecular data (pers. obs.) show that these two species are not directly related. A population of plants on Deer Spring Point, Kane County, Utah, appears to be this species, but molecular data indicate that it is probably a hybrid between P. tumulosa and, most likely, P. intermedia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 651. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella navajoensis |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (O’Kane) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) |
Web links |