Physaria chambersii |
Physaria multiceps |
|
---|---|---|
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
manyhead bladderpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex (buried), branched, (not thickened); densely pubescent, trichomes (sessile or short-stalked), 5–8-rayed, rays furcate or bifurcate, (umbonate, rough to finely tuberculate throughout). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
several from base, prostrate, (slender, sparsely pubescent), 0.5–2 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade obovate to narrowly elliptic, 1.5–6 cm, margins usually entire, rarely shallowly dentate, (surfaces densely pubescent, often silvery). |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
blade oblanceolate to spatulate, 0.5–1 cm, margins entire, (surfaces often sparsely pubescent). |
Racemes | congested. |
(narrow), loose, (elongated in fruit). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (greenish brown, sometimes magenta), linear or elliptic, 4.3–6(–7.5) mm, (median pair thickened apically, cucullate); petals (frequently pink or magenta in distal 1/3–1/2), spatulate to oblanceolate, 6–10(–12) mm, (claw undifferentiated from blade). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(ascending to somewhat spreading, straight to slightly curved), 4–8(–12) mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
broadly ovoid to suborbicular, inflated, (terete or, often, slightly angustiseptate), 3–6mm; valves sparsely pubescent; ovules usually 4, rarely 6–8 per ovary; style 3–6.5 mm. |
Seeds | flattened. |
plump. |
2n | = 8, 10, 16, 24. |
|
Physaria chambersii |
Physaria multiceps |
|
Phenology | Flowering Apr–Jul. | Flowering May–Jul. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Douglas-fir or spruce woodlands, limestone ridges, damp open slopes, soil pockets among rocks, crevices of rocks, decomposed calcareous rocks |
Elevation | 1500-3200 m (4900-10500 ft) | 2400-2900 m (7900-9500 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
|
ID; UT; WY |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 651. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella multiceps |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (Maguire) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) |
Web links |