Physaria chambersii |
Physaria montana |
|
---|---|---|
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
mountain bladderpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex simple or branched, (often enlarged); densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays furcate or bifurcate, (tuberculate throughout). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
simple or several from base, prostrate to erect, 0.5–2(–3.5) dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade suborbicular or obovate to elliptic, (1–)2–5(–7) cm, margins entire, sinuate, or shallowly dentate. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(often secund, proximal shortly petiolate, distal sessile); blade linear to obovate or rhombic, 1–2.5(–4) cm, margins entire or shallowly dentate. |
Racemes | congested. |
dense, compact, (usually elongated in fruit). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals elliptic, 5–8.5 mm, (lateral pair boat-shaped, saccate, median pair thickened apically, cucullate); petals (yellow to orange, sometimes fading purplish), narrowly spatulate or obovate, (6–)7.5–12 mm, (claw undifferentiated from blade, or gradually narrowed to claw, slightly expanded basally). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(usually sharply sigmoid, rarely nearly divaricate-spreading and straight), 5–15(–20) mm, (stout). |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(erect), ellipsoid or ovoid, not or slightly obcompressed, (apex not compressed), (6–)7–12 mm; valves densely pubescent, sometimes sparsely pubescent inside; ovules (8–)12–20(–24) per ovary; style 3–7 mm, (sometimes pubescent). |
Seeds | flattened. |
flattened. |
2n | = 8, 10, 16, 24. |
= 10. |
Physaria chambersii |
Physaria montana |
|
Phenology | Flowering Apr–Jul. | Flowering Apr–Jun(-Aug). |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Banks, rock outcrops, stony slopes and benchlands, from plains into mountains, in sagebrush, open scrub oak, pinyon-juniper woodland, ponderosa pine, Douglas fir on granitic, often gravelly, non-calcareous soils, rarely on calcareous soils |
Elevation | 1500-3200 m (4900-10500 ft) | 1000-3300 m (3300-10800 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
|
AZ; CO; NE; NM; SD; WY
|
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria montana is a rather variable species that in southwestern Colorado morphologically approaches P. rectipes and in eastern Wyoming approaches P. curvipes; it is unusual in the genus for its frequent presence on igneous, non-calcareous soils. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 650. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Vesicaria montana, Alyssum grayanum, Lesquerella montana, Lesquerella montana var. suffruticosa, Lesquerella rosulata |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (A. Gray) Greene: Fl. Francisc., 249. (1891) |
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