Physaria chambersii |
Physaria mcvaughiana |
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Mcvaugh's bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex simple or branched, (sometimes enlarged); densely pubescent, trichomes (sessile), several-rayed, rays fused (webbed) most of their length, (umbonate, peltate, tuberculate throughout). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
few to several from base, erect or outer ones decumbent, 0.5–4 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
(long-petiolate); blade elliptic to obovate or rhombic, 2–6(–9) cm, margins entire. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(sessile or shortly petiolate); blade oblanceolate to spatulate, 1–3 cm, (proximal broader), margins entire. |
Racemes | congested. |
dense, (relatively short). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals elliptic or narrowly oblong, 4–5.4 mm, (tapered to apex); petals (white, base and claw yellow, conspicuously purple-veined), usually broadly obovate or rhombic, 6–10 mm, (± equal to blade, tapering to slender claw). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(erect to spreading, ascending, or (proximal) horizontal, straight to slightly curved, sometimes loosely sigmoid), 6–12(–20) mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(sessile or substipitate, often reddish magenta), usually ovoid to subglobose, inflated, 4–6(–7) mm; valves (not retaining seeds after dehiscence), glabrous; replum as wide as or wider than fruit; septum perforate; ovules 8–12 per ovary; style 1.5–4 mm. |
Seeds | flattened. |
somewhat flattened. |
2n | = 8, 10, 16, 24. |
= 12. |
Physaria chambersii |
Physaria mcvaughiana |
|
Phenology | Flowering Apr–Jul. | Flowering mid Mar–Apr(-Aug). |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Stream bed gravels, rocky limestone slopes and hills, canyon bottoms and slopes, limestone rubble |
Elevation | 1500-3200 m (4900-10500 ft) | 1200-1600 m (3900-5200 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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TX; Mexico (Coahuila) |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 650. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella mcvaughiana |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) |
Web links |