Physaria chambersii |
Physaria lindheimeri |
|
---|---|---|
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Lindheimer's bladderpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Annuals or biennials; with a fine taproot; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate at base, sometimes bifurcate, (rough-tuberculate throughout). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
several from base, erect or outer decumbent, (often several-branched, branches slender and flexuous), to 8 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade pinnatisect to repand, 3–9(–14) cm, margins entire. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(sometimes secund, proximal usually petiolate, distal sessile); blade elliptic, 1–6 cm, (distal with cuneate base), margins entire or deeply dentate. |
Racemes | congested. |
dense. |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals elliptic to oblong, 3–5.5 mm, (median pair slightly thickened apically, cucullate); petals (sometimes drying slightly purplish), suborbicular or broadly ovate, 4.5–7(–9) mm, (narrowing gradually to short claw). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(horizontal or recurved and ascending at tip, sometimes loosely sigmoid), (5–)10–20 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
globose or broadly ellipsoid, not or slightly inflated, (4–)5–8 mm, (smooth); valves (not retaining seeds after dehiscence), glabrous; replum as wide as or wider than fruit; ovules (8–)12–16(–20) per ovary; style (1.5–)2–3(–4) mm. |
Seeds | flattened. |
flattened. |
2n | = 8, 10, 16, 24. |
= 12. |
Physaria chambersii |
Physaria lindheimeri |
|
Phenology | Flowering Apr–Jul. | Flowering Dec–Apr. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Heavy, black, claylike soils, or lighter, sandy soils, thickets, field-margins, roadsides, coastal prairies |
Elevation | 1500-3200 m (4900-10500 ft) | 20-800 m (100-2600 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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TX; Mexico (Tamaulipas) |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 649. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Vesicaria lindheimeri, Alyssum lindheimeri, Lesquerella gracilis var. pilosa, Lesquerella lindheimeri |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) |
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