Physaria chambersii |
Physaria lesicii |
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Pryor Mountains bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; (delicate, short-lived); caudex simple, (sometimes elongated, covered with persistent leaf bases); usually sparsely pubescent, trichomes 7–12-rayed, rays furcate near base. |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
simple from base, erect to decumbent, (unbranched, mostly filiform, slender), 1–1.5 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
(erect, petiole slender); blades broadly ovate to elliptic, 0.5–1 cm, (base abruptly narrowing to petiole), margins entire. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(remote, distally shortly petiolate); blade ± spatulate, (base cuneate), margins entire. |
Racemes | congested. |
lax, (elongated, few-flowered). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (erect), oblong, 3.5–4 mm, (lateral pair not saccate); petals (often fading to light purple apically), spatulate to nearly lingulate, 6–7 mm. |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(recurved to widely spreading, filiform, slender), 5–10 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(pendent), globose or subglobose, compressed, 3–4 mm; valves ± densely pubescent; ovules 6–10 per ovary; style ca. 1.5 mm. |
Seeds | flattened. |
not seen. |
2n | = 8, 10, 16, 24. |
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Physaria chambersii |
Physaria lesicii |
|
Phenology | Flowering Apr–Jul. | Flowering Jun(-early Jul). |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Pryor Mountains, on limestone soils in woodlands of Rocky Mountain juniper and/or mountain mahogany, and widely scattered Douglas-fir, fellfields dominated by bluebunch wheatgrass and cushion plants |
Elevation | 1500-3200 m (4900-10500 ft) | 1600-2000 m (5200-6600 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
|
MT |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 649. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella lesicii |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) |
Web links |