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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod

Kane County twinpod

Habit Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). Perennials; caudex simple, (with deep roots, thickened); densely (silvery) pubescent throughout (densely covering leaves with several appressed layers), less dense on stems, trichomes (stellate-scalelike), rays fused (webbed) in proximal 1/2 or to tips, (umbonate, nearly smooth to moderately tuberculate).
Stems

several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm.

simple from base, erect or outer ones slightly decumbent toward base, (from below or in basal leaves, unbranched), (0.5–)0.8–1.6(–2) dm.

Basal leaves

(petiole slender);

blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate.

(erect, petiole long, slender);

blade spatulate to broadly oblanceolate, (3–)5–7(–12) cm, (base gradually tapering to petiole), margins entire, (apex rounded or obtuse).

Cauline leaves

blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute).

blade oblanceolate, similar to basal, (base cuneate), margins entire.

Racemes

congested.

dense.

Flowers

sepals narrowly lanceolate, 5–8(–9) mm;

petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade).

sepals (erect), linear to linear-oblong, somewhat boat-shaped, 7–10 mm;

petals (erect at anthesis), lingulate, 11–15 mm, (claw undifferentiated from blade).

Fruiting pedicels

(divaricate, slightly sigmoid), 8–15 mm.

(divaricate-ascending, straight or slightly curved), 10–15 mm.

Fruits

(often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded);

valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent;

replum oblong, as wide as or wider than fruits, apex obtuse;

ovules 4–12 per ovary;

style (4–)6–8 mm (exceeding sinus).

(purplish in age), strongly didymous, semiorbicular, highly inflated, 10–18 × 14–19 mm, (papery), basal sinus usually shallow, rarely absent, apical sinus deep, narrowly V-shaped;

valves retaining seeds after dehiscence, sides flat, back rounded, margins keeled, base and apex obtuse;

replum narrowly oblong to linear, as wide as or wider than fruit, base slightly narrowed, apex obtusely rounded;

ovules 4 per ovary;

style 3–5 mm, (slender).

Seeds

flattened.

slightly flattened.

2n

= 8, 10, 16, 24.

Physaria chambersii

Physaria lepidota

Phenology Flowering Apr–Jul.
Habitat Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas
Elevation 1500-3200 m (4900-10500 ft)
Distribution
from FNA
AZ; CA; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
UT
[BONAP county map]
Discussion

Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Trichomes: rays fused nearly to tips; fruits with deep sinuses, or shallow basally, deep apically.
subsp. lepidota
1. Trichomes: rays fused in proximal 1/2; fruits with sinuses absent or shallow basally, deep apically.
subsp. membranacea
Source FNA vol. 7, p. 631. FNA vol. 7, p. 648.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. lepidota subsp. lepidota, P. lepidota subsp. membranacea
Synonyms P. chambersii var. canaani, P. chambersii var. sobolifera
Name authority Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) Rollins: Brittonia 33: 335, figs. 1, 2. (1981)
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