Physaria chambersii |
Physaria klausii |
|
---|---|---|
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Rogers Pass or klaus' or divide bladderpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex simple; densely pubescent, trichomes (loosely spreading), 3–5-rayed, rays distinct, furcate (with exceptionally long branches). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
simple from base, erect to decumbent, (slender), 0.6–1.5 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blades obovate to deltate, 1.5–3(–4) cm, margins entire or outer one with 1 or 2 broad teeth. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
blade oblanceolate to spatulate, 0.6–1.5 cm, margins entire. |
Racemes | congested. |
loose. |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (green-yellow, often tinged with purple), elliptic, 3–4.6 mm; petals oblanceolate, 6–8 mm (claw expanded). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(sigmoid), 5–9 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(depressed), broadly obovate, compressed (angustiseptate), 2–4 mm, (apex slightly bilobed to nearly truncate); valves densely pubescent, trichomes strongly ascending, spreading, long, (appearing fuzzy), pubescent inside; ovules 4 per ovary; style 3–4 mm, (pubescent or glabrous). |
Seeds | flattened. |
flattened. |
2n | = 8, 10, 16, 24. |
|
Physaria chambersii |
Physaria klausii |
|
Phenology | Flowering Apr–Jul. | Flowering Jul. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Open gravel slides, solifluction cross-stripes of shale rubble, barren shale-derived soil |
Elevation | 1500-3200 m (4900-10500 ft) | 1200-1900 m (3900-6200 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
|
MT |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 647. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella klausii |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) |
Web links |