Physaria chambersii |
Physaria integrifolia |
|
---|---|---|
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Snake River or creeping twinpod, Snake River twinpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex usually branched, (rhizomelike, cespitose); densely (silvery) pubescent, trichomes (often stalked, appressed), several-rayed, rays furcate or bifurcate, (umbonate, strongly tuberculate throughout). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
several from base, ± erect, exceeding basal rosette by ± 0.5 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
(forming a strong rosette; long-petiolate); blade oblanceolate to ovate or orbicular, (1.5–)2–4(–8) cm, (base usually abruptly tapering to petiole), margins entire. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
blade oblanceolate, 1–2 cm, margin entire, (apex acute). |
Racemes | congested. |
congested, (greatly exceeding leaves). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals often keeled, 6–8 mm; petals spatulate, 8–10 mm. |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(spreading, straight or slightly curved), 7–11 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
didymous, highly inflated, 8–22 × 10–25 mm, (papery, basal and apical sinuses deep); valves (retaining seeds after dehiscence), densely pubescent, trichomes appressed; replum linear to oblong, as wide as or wider than fruit; ovules 8 per ovary; style 7–9 mm. |
Seeds | flattened. |
flattened. |
2n | = 8, 10, 16, 24. |
= 16. |
Physaria chambersii |
Physaria integrifolia |
|
Phenology | Flowering Apr–Jul. | Flowering (May-)Jun–Jul(-Aug). |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Calcareous hills and slopes, shale-limestone cliffs, bare steep slopes, red clay banks, shale |
Elevation | 1500-3200 m (4900-10500 ft) | 1900-2700 m (6200-8900 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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ID; MT; WY |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria integrifolia has traditionally been recognized as a variety of P. didymocarpa, but it is morphologically and ecologically quite distinctive. Variety monticola (no combination has been made at subspecific rank) is not recognized here; it is considered another example, in the genus, of caudices elongating in response to shifting substrates. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 644. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | P. didymocarpa var. integrifolia, P. integrifolia var. monticola |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (Rollins) Lichvar: Madroño 31: 203. (1984) |
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