Physaria chambersii |
Physaria humilis |
|
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
St. Mary's Peak or bitterroot bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex simple, (thickened with persistent leaf bases); densely (silvery) pubescent, trichomes 5- or 6-rayed, rays furcate or 3-branched. |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
simple or few to several from base, prostrate, (from below a terminal rosette of leaves, unbranched), 0.2–0.5 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade elliptic to broadly ovate or obovate, (1–)1.5–2.5(–3) cm, margins entire, (apex obtuse). |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
blade spatulate, 3–7 mm, (base cuneate), margins entire. |
Racemes | congested. |
not loose, (scarcely elongated in fruit, 3–5-flowered). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (yellow-green), narrowly elliptic to narrowly long-triangular, 3.7–5 mm; petals oblanceolate to nearly obovate, 7–8.5 mm, (abruptly tapering to narrow claw). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(straight or slightly curved). |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
wider than long, apex truncate to shallowly notched, compressed (angustiseptate), 3–4 mm; valves densely pubescent, trichomes ascending to erect, sparsely pubescent inside; ovules 4 per ovary; style 2–3 mm. |
Seeds | flattened. |
plump, (slightly compressed). |
2n | = 8, 10, 16, 24. |
|
Physaria chambersii |
Physaria humilis |
|
Phenology | Flowering Apr–Jul. | Flowering Jun-early Aug. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Steep slopes, dry summits, rocky fellfields, dry ledges |
Elevation | 1500-3200 m (4900-10500 ft) | 2700-2900 m (8900-9500 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
|
MT |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria humilis is found in metamorphosed rock and detritus on the peaks of the Bitterroot Mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 644. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella humilis |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) |
Web links |