Physaria chambersii |
Physaria hemiphysaria |
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
intermountain bladderpod, skyline bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex simple or branched (tightly); sparsely to densely pubescent, trichomes (sessile or short-stalked), 4–6-rayed, rays furcate or bifurcate, distinct or slightly fused at base, (sometimes umbonate, tuberculate throughout). | ||||
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
few to several from base, decumbent, 0.5–1(–2) dm, (rather stout, sparsely pubescent). |
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Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade elliptic to suborbicular, 1.5–3.5(–5.5) cm, margins entire or shallowly dentate (at base, surfaces densely pubescent, silvery). |
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Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(petiolate or distal nearly sessile); blade elliptic to obovate, 0.5–1.5 cm, margins entire. |
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Racemes | congested. |
dense, congested, (few-flowered). |
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Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals lanceolate, oblanceolate, or narrowly elliptic, 3.8–5 mm, (median pair thickened apically, cucullate); petals narrowly lanceolate to linear, 6–10(–13) mm. |
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Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(spreading or recurved, sometimes loosely sigmoid), 2–6.5 mm. |
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Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(sessile or substipitate), broadly obcordate, obdeltate, or obcompressed, slightly compressed (angustiseptate), 3–5(–7) mm; valves (not retaining seeds after dehiscence), sparsely pubescent or glabrous, trichomes appressed; replum as wide as or wider than fruit; ovules 8–16 per ovary; style (1.8–)3–6(–7) mm. |
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Seeds | flattened. |
slightly flattened, (ellipsoid to suborbicular). |
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2n | = 8, 10, 16, 24. |
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Physaria chambersii |
Physaria hemiphysaria |
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Phenology | Flowering Apr–Jul. | |||||
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | |||||
Elevation | 1500-3200 m (4900-10500 ft) | |||||
Distribution |
AZ; CA; ID; NV; OR; UT
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UT |
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 642. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella hemiphysaria | ||||
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (Maguire) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | ||||
Web links |