Physaria chambersii |
Physaria grahamii |
|
---|---|---|
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Graham's twinpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex branched, (thick, cespitose); densely pubescent, trichomes rays (appressed on leaves, ascending on pedicels and fruits), distinct, furcate or bifurcate. |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
several from base, decumbent to erect or ascending (unbranched), 1–2.5 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
(outer ones spreading, inner erect or ascending); blade ovate, often broadly so, 4–7 cm, margins repand to lyrate-lobed. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
similar to basal, blade oblanceolate or narrowly oblong, reduced in size, (base gibbous). |
Racemes | congested. |
loose, (elongated). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals lanceolate or narrowly oblong, 5.8–7.2 mm; petals (erect, sometime purplish or drying purple), narrowly oblong to oblanceolate, 7–10 mm, (not or weakly clawed). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(ascending to divaricate-ascending, sigmoid to nearly straight), 10–17 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
didymous, globose or subglobose, inflated, 10–13 mm, (papery, basal and apical sinuses deep); valves (retaining seeds after dehiscence), pubescent, trichomes ascending, appearing fuzzy; replum oblong to oblanceolate, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style (4–)5–7 mm. |
Seeds | flattened. |
plump, (suborbicular). |
2n | = 8, 10, 16, 24. |
|
Physaria chambersii |
Physaria grahamii |
|
Phenology | Flowering Apr–Jul. | Flowering May–Jun. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Sagebrush, pinyon-juniper, ponderosa pine, Douglas-fir, limber pine communities on clay, or a mixture of shale fragments and clay |
Elevation | 1500-3200 m (4900-10500 ft) | 2100-2900 m (6900-9500 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
|
UT |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria grahamii is difficult to evaluate due to the paucity of collections. The tentative recognition by N. H. Holmgren (2005b) is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 642. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | P. acutifolia var. purpurea, P. acutifolia var. repanda, P. repanda |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | C. V. Morton: Ann. Carnegie Mus. 26: 220. (1937) |
Web links |