The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod

Goodding's bladderpod

Habit Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). Annuals or biennials; without caudex, (cespitose); densely pubescent, trichomes (sessile or short-stalked), few-rayed, rays (ascending or erect), simple or infrequently furcate near base, (long and slender, sometimes with U-shaped notch on one side, smooth or finely tuberculate).
Stems

several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm.

several from base, erect (and stout) or outer ones decumbent, (unbranched or branched, stiff and densely foliate, sterile leaf-bearing branches sometimes present), to 4 dm.

Basal leaves

(petiole slender);

blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate.

blade obovate or elliptic, to ca. 3 cm, margins sinuate or shallowly dentate.

Cauline leaves

blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute).

(proximal usually shortly petiolate, distal sessile);

blade obovate to broadly elliptic, 1–3 cm, margins sinuate or shallowly toothed.

Racemes

congested.

dense, compact, (elongated in fruit).

Flowers

sepals narrowly lanceolate, 5–8(–9) mm;

petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade).

sepals elliptic or narrowly elliptic or oblong, (3.8–)4.5–5.5 mm, (lateral pair cucullate, very convex, median pair tapering to base, thickened apically, cucullate, often slightly keeled);

petals cuneate, 6.5–8 mm, (slightly expanded at base, margins sinuate, apex retuse or entire).

Fruiting pedicels

(divaricate, slightly sigmoid), 8–15 mm.

(recurved, curved or sigmoid), somewhat expanded apically.

Fruits

(often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded);

valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent;

replum oblong, as wide as or wider than fruits, apex obtuse;

ovules 4–12 per ovary;

style (4–)6–8 mm (exceeding sinus).

(sessile or substipitate), oblong or broadly elliptic, compressed (latiseptate), 5–8 mm;

valves pubescent, trichomes spreading, sparsely pubescent inside;

ovules 4–6 per ovary;

style 3–5 mm.

Seeds

flattened.

flattened.

2n

= 8, 10, 16, 24.

Physaria chambersii

Physaria gooddingii

Phenology Flowering Apr–Jul. Flowering Jun–Sep.
Habitat Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas Mountainous areas, open areas in pinyon-juniper and ponderosa pine forests
Elevation 1500-3200 m (4900-10500 ft) 1800-2300 m (5900-7500 ft)
Distribution
from FNA
AZ; CA; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM
[BONAP county map]
Discussion

Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Physaria gooddingii (found in the mountains of Catron, Sierra, and western Socorro counties, New Mexico, and in Greenlee County, Arizona) is similar to 9. P. aurea (found farther east), but differs in having trichomes with ascending or erect rays (rather than appressed) and fruits that are strongly latiseptate (rather than not, or very little, compressed), a state that is infrequent in the genus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 631. FNA vol. 7, p. 639.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms P. chambersii var. canaani, P. chambersii var. sobolifera Lesquerella gooddingii
Name authority Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) (Rollins & E. A. Shaw) O’Kane & Al-Shehbaz: Novon 12: 323. (2002)
Web links