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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod

globe bladderpod

Habit Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). Biennials or perennials; caudex branched, (± woody); densely pubescent, trichomes (sessile), 3–6-rayed, rays distinct and simple or furcate, (in 2 layers, lower layer umbonate, smooth to finely tuberculate, some often with a U-shaped notch).
Stems

several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm.

several from base, erect, (arising among leaves of an elongated main axis), to 5 dm (± equal).

Basal leaves

(petiole slender);

blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate.

(shortly petiolate);

blade obovate to oblanceolate, (1.5–)2.5–5(–6) cm, margins entire, sinuate to shallowly toothed, or pinnatifid.

Cauline leaves

blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute).

(sessile or shortly petiolate);

blade oblanceolate to oblong, 1.3–3(–4) cm, (base cuneate), margins entire or repand to dentate.

Racemes

congested.

dense.

Flowers

sepals narrowly lanceolate, 5–8(–9) mm;

petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade).

sepals elliptic or obovate, 2.6–4.1 mm, (median pair thickened apically, cucullate);

petals (bright yellow), obovate, 3.5–6.5(–7.5) mm, (margins sinuate).

Fruiting pedicels

(divaricate, slightly sigmoid), 8–15 mm.

(usually spreading horizontally, straight), 7–14(–21) mm.

Fruits

(often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded);

valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent;

replum oblong, as wide as or wider than fruits, apex obtuse;

ovules 4–12 per ovary;

style (4–)6–8 mm (exceeding sinus).

(sessile or substipitate);

globose, often slightly compressed apically, (1–)2–3 mm;

valves sparsely pubescent, sometimes pubescent inside, trichomes spreading, 3–5-rayed;

ovules 4 per ovary;

style 2–3.5(–4) mm.

Seeds

flattened.

flattened or plump, (often outer surface hemispherical, inner surface flattened, or both surfaces rounded).

2n

= 8, 10, 16, 24.

= 14.

Physaria chambersii

Physaria globosa

Phenology Flowering Apr–Jul. Flowering Mar–May.
Habitat Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas Open rocky areas, shale at cliff bases, open talus, ledges, open cedar glades
Elevation 1500-3200 m (4900-10500 ft) 100-300 m (300-1000 ft)
Distribution
from FNA
AZ; CA; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
IN; KY; TN
[BONAP county map]
Discussion

Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Physaria globosa is possibly introduced in Indiana. A report for Ohio was based on a collection by “Jones,” but that specimen cannot be located.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 631. FNA vol. 7, p. 639.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms P. chambersii var. canaani, P. chambersii var. sobolifera Vesicaria globosa, Alyssum globosum, Alyssum shortii, Lesquerella globosa
Name authority Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) (Desvaux) O’Kane & Al-Shehbaz: Novon 12: 323. (2002)
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