Physaria chambersii |
Physaria fremontii |
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Fremont's bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex simple; densely pubescent, trichomes (subsessile), 5–7-rayed, rays distinct or slightly fused at base, usually furcate, (roughly tuberculate). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
few to several from base, prostrate, (arising proximal to a terminal cluster of erect leaves, usually unbranched, slender), 0.5–1(–1.5) dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
(petiole slender); blade elliptic to rhombic, 1.5–4(–5) cm, (base gradually tapering to petiole), margins usually entire, rarely dentate. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
blade narrowly obovate to oblanceolate, 5–15 mm, margins entire. |
Racemes | congested. |
(relatively short), rather loose. |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals ± elliptic, 4–6 mm; petals ovate, 6–8 mm, (claw slightly expanded at base). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(usually secund, recurved), 5–8 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(pendent), globose, subglobose, or slightly obcompressed, slightly or not inflated, 3–6 mm, (rigid, apex usually beaked); valves pubescent throughout; ovules 8–12 per ovary; style 1–2 mm, (pubescent). |
Seeds | flattened. |
slightly flattened, (suborbicular). |
2n | = 8, 10, 16, 24. |
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Physaria chambersii |
Physaria fremontii |
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Phenology | Flowering Apr–Jul. | Flowering May–Jun. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Calcareous gravel, loose whitish rubble, limestone pavement, rocky calcareous ridges |
Elevation | 1500-3200 m (4900-10500 ft) | 2100-2800 m (6900-9200 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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WY |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria fremontii is known from the area of the Wind River Mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 638. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella fremontii |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (Rollins & E. A. Shaw) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) |
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