Physaria chambersii |
Physaria floribunda |
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
point-tip twinpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays mostly furcate, (arms of unequal lengths, finely tuberculate). | ||||
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
several from base, erect or lateral decumbent, (unbranched), 1–2 dm. |
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Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
(petiole usually winged); blade broadly oblanceolate, 3–8 cm, margins usually dentate or pinnatifid, rarely subentire, (terminal lobe acute or obtuse, not rounded). |
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Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
blade spatulate to linear-oblanceolate, 1–3 cm, margins usually entire, rarely toothed, (apex acute). |
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Racemes | congested. |
loose (and greatly elongated in fruit) to congested. |
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Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals linear-oblong, 5–7 mm; petals spatulate, 9–11 mm. |
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Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(recurved), 6–15 mm. |
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Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(usually pendent on arching pedicels, less frequently widely divergent), irregular in shape, (base obtuse or slightly cordate, apex deeply and broadly notched), not strongly inflated, 8–11 × 8–12 mm, (papery); valves retaining seeds after dehiscence; replum linear-oblong, constricted, 2.5–4 mm, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style 5–8 mm. |
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Seeds | flattened. |
flattened. |
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2n | = 8, 10, 16, 24. |
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Physaria chambersii |
Physaria floribunda |
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Phenology | Flowering Apr–Jul. | |||||
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | |||||
Elevation | 1500-3200 m (4900-10500 ft) | |||||
Distribution |
AZ; CA; ID; NV; OR; UT
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CO; NM
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 637. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | |||||
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | ||||
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