Physaria chambersii |
Physaria filiformis |
|
---|---|---|
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
limestone glade or Missouri bladderpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Annuals; with a fine taproot; densely pubescent, trichomes (sessile), usually 4-rayed, rays forked, rarely simple or tripartite, (finely tuberculate). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
several from base, (slender), erect or outer decumbent, (usually branched, branches filiform, bud clusters of growing plants drooping), to 2.5 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade orbicular to broadly spatulate, 1–2.4 cm, margins entire or sinuate. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(proximal often petiolate, distal sessile); similar to basal, blade spatulate to oblanceolate or (distal) linear, (base cuneate), margins entire or sinuate. |
Racemes | congested. |
loose. |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals oblong or elliptic, 2.5–4.6 mm, (median pair slightly thickened apically); petals (pale yellow), spatulate to obovate, 5–9 mm, (apex ± emarginate). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(usually divaricate-ascending, straight or slightly curved), 7–11 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(sessile or shortly stipitate), globose, not inflated, 3–4 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4 per ovary; style 3–5 mm. |
Seeds | flattened. |
flattened. |
2n | = 8, 10, 16, 24. |
= 14. |
Physaria chambersii |
Physaria filiformis |
|
Phenology | Flowering Apr–Jul. | Flowering Apr–May. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Limestone, dolomite, and shale, sparsely vegetated or barren areas, cedar glades, old pastures, along roadsides |
Elevation | 1500-3200 m (4900-10500 ft) | 200-300 m (700-1000 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
|
AL; AR; MO |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 637. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella filiformis |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) |
Web links |