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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod

Engelmann's bladderpod

Habit Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). Perennials; caudex simple or branched, (woody, aerial); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, distinct or fused at base, (asymmetrical with deep notch on one side, often with a U-shaped gap between 2 of the rays, umbonate, strongly tuberculate).
Stems

several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm.

few to several from base, erect, (usually unbranched), (1.5–)2.5–4(–6) dm.

Basal leaves

(petiole slender);

blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate.

blade elliptic to obovate, 2–6.5 cm, margins entire, sinuate, or remotely toothed, (surfaces occasionally sparsely pubescent).

Cauline leaves

blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute).

(proximal often petiolate, distal sessile or subsessile);

blade oblanceolate to linear, 1–4 cm, margins entire.

Racemes

congested.

dense, (subumbellate).

Flowers

sepals narrowly lanceolate, 5–8(–9) mm;

petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade).

sepals ovate or elliptic, 5.5–10 mm, (median pair thickened apically, cucullate);

petals (bright yellow), obovate to elliptic, 8–14 mm, (sometimes with distinct claw, often retuse).

Fruiting pedicels

(divaricate, slightly sigmoid), 8–15 mm.

(ascending), relatively short.

Fruits

(often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded);

valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent;

replum oblong, as wide as or wider than fruits, apex obtuse;

ovules 4–12 per ovary;

style (4–)6–8 mm (exceeding sinus).

(shortly stipitate), ± globose or ellipsoid, not or slightly inflated, 5–8 mm;

valves (not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules (8–)12–20 per ovary;

style 3.5–5 mm.

Seeds

flattened.

flattened.

2n

= 8, 10, 16, 24.

= 12, 24, 36.

Physaria chambersii

Physaria engelmannii

Phenology Flowering Apr–Jul. Flowering Apr–May.
Habitat Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas Limestone prairies, rocky ridges, pebbly shores, thin caliche soils, limestone outcrops
Elevation 1500-3200 m (4900-10500 ft) 150-400 m (500-1300 ft)
Distribution
from FNA
AZ; CA; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
OK; TX
[BONAP county map]
Discussion

Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 631. FNA vol. 7, p. 635.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms P. chambersii var. canaani, P. chambersii var. sobolifera Vesicaria engelmannii, Alyssum engelmannii, Lesquerella engelmannii, Vesicaria engelmannii var. elatior, Vesicaria pulchella
Name authority Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 322. (2002)
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