Physaria chambersii |
Physaria engelmannii |
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Engelmann's bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex simple or branched, (woody, aerial); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, distinct or fused at base, (asymmetrical with deep notch on one side, often with a U-shaped gap between 2 of the rays, umbonate, strongly tuberculate). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
few to several from base, erect, (usually unbranched), (1.5–)2.5–4(–6) dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade elliptic to obovate, 2–6.5 cm, margins entire, sinuate, or remotely toothed, (surfaces occasionally sparsely pubescent). |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(proximal often petiolate, distal sessile or subsessile); blade oblanceolate to linear, 1–4 cm, margins entire. |
Racemes | congested. |
dense, (subumbellate). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals ovate or elliptic, 5.5–10 mm, (median pair thickened apically, cucullate); petals (bright yellow), obovate to elliptic, 8–14 mm, (sometimes with distinct claw, often retuse). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(ascending), relatively short. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(shortly stipitate), ± globose or ellipsoid, not or slightly inflated, 5–8 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules (8–)12–20 per ovary; style 3.5–5 mm. |
Seeds | flattened. |
flattened. |
2n | = 8, 10, 16, 24. |
= 12, 24, 36. |
Physaria chambersii |
Physaria engelmannii |
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Phenology | Flowering Apr–Jul. | Flowering Apr–May. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Limestone prairies, rocky ridges, pebbly shores, thin caliche soils, limestone outcrops |
Elevation | 1500-3200 m (4900-10500 ft) | 150-400 m (500-1300 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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OK; TX |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 635. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Vesicaria engelmannii, Alyssum engelmannii, Lesquerella engelmannii, Vesicaria engelmannii var. elatior, Vesicaria pulchella |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) |
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