Physaria chambersii |
Physaria eburniflora |
|
---|---|---|
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Devils Gate twinpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex usually simple; densely pubescent, trichomes (sessile), rays often furcate, fused toward base, (nearly smooth). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
simple from base, prostrate, (arising lateral to rosette), 0.1–0.5 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade suborbicular, (1–)2.5(–3) cm, (base abruptly narrowed to petiole), margins entire, (flat), (surfaces densely silvery pubescent, trichomes in multiple layers, appressed). |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(2–4); blade oblanceolate, ca. 1 cm, (base cuneate), margins entire, (apex acute). |
Racemes | congested. |
condensed. |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (erect, purplish to greenish), linear-oblong or boat-shaped, 5.5–6.5 mm, (lateral pair more saccate than median); petals (white), spatulate, 9–12 mm, (claw undifferentiated from blade). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(divaricate-ascending, nearly straight), 6–10 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
strongly didymous, irregular in shape and size, (base slightly cordate, apex with a deep closed sinus), strongly to somewhat inflated, 6–8 × 6–8 mm (± bladderlike, papery); valves (retaining seeds after dehiscence), pubescent; replum elliptic to obovate, not constricted, as wide as or wider than fruit, apex obtuse; ovules 4–8 per ovary; style 4–5 mm, (sparsely pubescent or glabrous). |
Seeds | flattened. |
plump. |
2n | = 8, 10, 16, 24. |
|
Physaria chambersii |
Physaria eburniflora |
|
Phenology | Flowering Apr–Jul. | Flowering May–Jun. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Limestone hills, red soil, rocky calcareous slopes, clay depressions, granite and marble detritus |
Elevation | 1500-3200 m (4900-10500 ft) | 1800-3000 m (5900-9800 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
|
WY |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 635. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | Rollins: Brittonia 33: 333. (1981) |
Web links |