Physaria chambersii |
Physaria douglasii |
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Columbia bladder-pod, Douglas' bladder-pod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). | ||||
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
simple from base, erect, (usually unbranched), to 4.5 dm. |
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Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid. |
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Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
similar to basal, blade narrowly linear or, sometimes, orbicular. |
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Racemes | congested. |
loose (lax). |
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Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate); petals 6–11 mm. |
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Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(recurved, straight, curved, or sigmoid), 6–20 mm. |
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Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm; valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked; ovules 4(–8) per ovary; style (1.6–)3–6 mm. |
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Seeds | flattened. |
flattened. |
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2n | = 8, 10, 16, 24. |
= 10, 30. |
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Physaria chambersii |
Physaria douglasii |
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Phenology | Flowering Apr–Jul. | |||||
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | |||||
Elevation | 1500-3200 m (4900-10500 ft) | |||||
Distribution |
AZ; CA; ID; NV; OR; UT
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ID; MT; OR; WA; BC
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 634. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella douglasii | ||||
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | ||||
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