Physaria chambersii |
Physaria dornii |
|
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Dorn's twinpod, tunp range twinpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; (compact); caudex simple, (stout); densely (silvery) pubescent throughout (except style), trichomes several-rayed, rays furcate, fused at base, (umbonate, tuberculate throughout). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
simple from base, erect, (arising from a condensed rosette), to 1 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
(ascending or erect); blade elliptic to oblanceolate to obovate, (usually curled from middle to apex), (1.5–)5–7 cm (width 12–20 mm), margins entire. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(1–5); blade oblanceolate, 1–2.5 cm, margins entire. |
Racemes | congested. |
compact (or elongated in fruit, to 1 dm, barely exceeding leaves). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (erect), oblong to linear or spatulate, 5.5–7 mm; petals spatulate, 10–14 mm, (claw undifferentiated from blade). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(divaricate-ascending, slightly curved), 7–18 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
didymous, irregular, highly inflated, 8–11(–18) × 10–15 mm, (papery, basal sinus shallower than the deep apical sinus; valves retaining seeds after dehiscence); replum obovate, not constricted, 1–1.8 mm, apex obtuse, as wide as or wider than fruit; ovules (4–)8(–12) per ovary; style 4–6 mm, (glabrous). |
Seeds | flattened. |
flattened, (oblong to elliptic, thin-margined or not). |
2n | = 8, 10, 16, 24. |
|
Physaria chambersii |
Physaria dornii |
|
Phenology | Flowering Apr–Jul. | Flowering May–Jun. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Calcareous shale, slopes, ridges |
Elevation | 1500-3200 m (4900-10500 ft) | 1900-2200 m (6200-7200 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
|
WY |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 634. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | Lichvar: Brittonia 35: 150, figs. 1–3. (1983) |
Web links |