Physaria chambersii |
Physaria densiflora |
|
---|---|---|
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
denseflower bladderpod, low bladderpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Annuals or biennials; caudex simple or branched, (relatively small, cespitose); densely pubescent, trichomes (spreading, sessile or short-stalked), 5–7-rayed, rays distinct and simple, (tuberculate, finely tubercled with a U-shaped notch on one side). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
simple or few to several from base, erect or decumbent, (rarely branched, usually leafy), to 4 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade lyrate-pinnatifid, 1–7 cm, margins entire or shallowly dentate. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(sessile or shortly petiolate); blade narrowly obovate to elliptic, 0.5–6 cm, margins entire, repand, or shallowly dentate. |
Racemes | congested. |
dense, (elongated in fruit, often subtended by distal cauline leaves). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals elliptic, 3.7–7.2 mm, (lateral pair somewhat cucullate, median pair thickened apically); petals (yellow to orange-yellow), obovate to obdeltate, (4.5–)7–10(–11) mm, (tapering to short claw, apex often emarginate). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(usually divaricate-spreading, straight or slightly curved, delicate, sometimes drooping, especially on herbarium specimens), 7–10 mm, (somewhat rigid). |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(sessile or substipitate), globose or broadly obovate, not inflated, 4–6 mm, (smooth); valves (not retaining seeds after dehiscence), glabrous; replum as wide as or wider than fruit; ovules 8–16 per ovary; style 2–5 mm. |
Seeds | flattened. |
flattened. |
2n | = 8, 10, 16, 24. |
= 14. |
Physaria chambersii |
Physaria densiflora |
|
Phenology | Flowering Apr–Jul. | Flowering Mar–May. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Sandy, granitic, or calcareous soils, sandy ledges, limestone outcrops, rocky prairies, uplands |
Elevation | 1500-3200 m (4900-10500 ft) | 30-400 m (100-1300 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
|
TX |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Alyssum densiflorum (A. Gray) Kuntze (1891), not Desfontaines (1808) is an illegitimate name, sometimes found in synonymy with Physaria densiflora. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 633. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Vesicaria densiflora, Lesquerella densiflora |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) |
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