Physaria chambersii |
Physaria congesta |
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Dudley bluffs bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; (relatively diminutive, strongly condensed); caudex (buried), simple or branched, (stout, thatched, thickened with persistent leaf bases); densely pubescent, trichomes (appressed, stiff), 4- or 5-rayed, rays fused at center, (mostly bifurcate). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
simple or few from base, decumbent to ascending, (arising laterally from a tight hemispherical tuft of leaves), to 0.15 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
similar to cauline, (erect, surfaces silvery). |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(ascending, subsessile); blade linear-oblanceolate, (0.6–)0.8–1.3(–1.5) cm, margins entire, (apex acute to narrowly obtuse). |
Racemes | congested. |
strongly congested, (often sessile or nearly so, lateral to leaves). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (loosely erect), narrowly oblong, 3–4 mm; petals spatulate, 5–6 mm. |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(erect or ascending, straight to slightly curved), 3–6 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
ovate, compressed (latiseptate) on margins and apically, 4–5 mm; valves pubescent, trichomes densely appressed; ovules 4 per ovary; style 1–1.5 mm. |
Seeds | flattened. |
plump. |
2n | = 8, 10, 16, 24. |
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Physaria chambersii |
Physaria congesta |
|
Phenology | Flowering Apr–Jul. | Flowering Apr–May. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Barren knolls with pinyon-juniper |
Elevation | 1500-3200 m (4900-10500 ft) | 1800-2100 m (5900-6900 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
|
CO |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria congesta is found on white, decomposed shale of the Thirteenmile Creek Tongue of the Green River Formation. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 632. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella congesta |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) |
Web links |