Physaria chambersii |
Physaria cinerea |
|
---|---|---|
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
basin bladderpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex branched, (woody); densely pubescent, trichomes (appressed, except spreading on pedicels and fruits, sessile or short-stalked), several-rayed, rays furcate or bifurcate, (strongly tuberculate throughout). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
few from base, erect, (stout), to 1.5 dm (sometimes greatly reduced). |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade suborbicular to rhombic or broadly elliptic, 1–4.5 cm, margins entire, (apex rounded to subacute). |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(proximal petiolate, distal subsessile); blade elliptic, 1–4 cm, (distal narrower), margins entire or remotely dentate. |
Racemes | congested. |
condensed, (subcorymbose to subumbellate, few-flowered). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (persistent), broadly ovate or oblong to narrowly elliptic, 5.5–8(–9.5) mm, (lateral pair slightly cucullate, median pair thickened apically, cucullate, usually keeled); petals (orange to yellow), oblong to obovate, 8–11.5(–14.5) mm, (slightly narrowed to broad claw, margins sinuate, often retuse). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(horizontal or divaricate-ascending, straight or slightly curved), 5–15 mm, (stout and rigid). |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(sessile or substipitate), globose, ovoid, or suborbicular, compressed (with marginal and apical constriction), 4–7 mm; valves pubescent, trichomes contiguous or overlapping, often spreading; ovules 16–24 per ovary; style 2–4 mm. |
Seeds | flattened. |
lenticular, ovate in outline. |
2n | = 8, 10, 16, 24. |
= 10. |
Physaria chambersii |
Physaria cinerea |
|
Phenology | Flowering Apr–Jul. | Flowering Mar–May. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Red soil, chiprock, gypsum or chalky knolls, limestone rubble |
Elevation | 1500-3200 m (4900-10500 ft) | 900-2200 m (3000-7200 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
|
AZ |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 631. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella cinerea |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) |
Web links |