Physaria chambersii |
Physaria carinata |
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Idaho bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex simple, (often enlarged by persistent leaf bases); densely pubescent, trichomes (sessile or short-stalked), rays furcate or bifurcate, (nearly smooth to finely tuberculate). | ||||||||
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
simple from base, decumbent, (occasionally few-branched), 0.5–1.5(–2) dm. |
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Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade elliptic to broadly obovate, triangular, rhombic, or round, 1.5–3(–4) cm, margins often sinuate or shallowly lobed. |
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Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(sessile or shortly petiolate); blade elliptic to oblanceolate to obovate, 0.5–1.5 cm, (base narrowed to petiole), margins entire. |
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Racemes | congested. |
compact to loose. |
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Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals oblong to broadly elliptic, 4–7.5 mm, (lateral pair saccate or not); petals spatulate, 7.5–10 mm. |
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Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(ascending or divaricate-spreading, straight to loosely sigmoid or curved), 4–10 mm. |
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Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(sessile or substipitate), elliptic, suborbicular, or oblong, strongly compressed (angustiseptate), 5–9 mm, (rounded to sharply keeled on 1 side, edges ± keeled); valves: (margins covering replum edges or not), usually pubescent throughout or, rarely, glabrous inside; ovules (4–)8–14(–16) per ovary; style 2–4.5(–5) mm. |
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Seeds | flattened. |
slightly flattened. |
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2n | = 8, 10, 16, 24. |
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Physaria chambersii |
Physaria carinata |
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Phenology | Flowering Apr–Jul. | |||||||||
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | |||||||||
Elevation | 1500-3200 m (4900-10500 ft) | |||||||||
Distribution |
AZ; CA; ID; NV; OR; UT
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ID; MT; WY |
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 3 (3 in the flora). Differences in fruit morphology become blurred and the three subspecies are often indistinguishable where their ranges meet near the intersection of Idaho, Montana, and Wyoming. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 629. | ||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella carinata | ||||||||
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | ||||||||
Web links |