Physaria chambersii |
Physaria calderi |
|
---|---|---|
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Calder's bladderpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex simple or branched; densely pubescent throughout, trichomes (sessile or subsessile), rays distinct or slightly fused at base, furcate or bifurcate, (strongly umbonate, tuberculate, tubercles often relatively larger, fewer over center). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
simple or few to several from base, usually erect to spreading, sometimes prostrate, 0.5–2 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade oblanceolate, 2–3 cm, margins entire. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(sessile or proximal shortly petiolate); blade narrowly oblanceolate, 0.5–1.5 cm, margins entire. |
Racemes | congested. |
loose. |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate); petals obovate, (6–)7–10 mm (nearly as wide, abruptly narrowed to claw, ca. 1 mm wide). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(erect to divaricate or ascending, sometimes curved), (5–)10–20(–40) mm, (stout). |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
subglobose to ellipsoid, compressed (usually angustiseptate), to 8 mm; (valves not retaining seeds after dehiscence); replum as wide as or wider than fruit; ovules 10–14 per ovary; style 1–2 mm. |
Seeds | flattened. |
plump. |
2n | = 8, 10, 16, 24. |
= 20. |
Physaria chambersii |
Physaria calderi |
|
Phenology | Flowering Apr–Jul. | Flowering Jun–Aug. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Dry rocky summits, limestone flats and slopes, alpine knolls |
Elevation | 1500-3200 m (4900-10500 ft) | 600-1500 m (2000-4900 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
|
AK; NT; YT |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria calderi is known from the Ogilvie and Richardson mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 629. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella calderi, Lesquerella arctica subsp. calderi |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (G. A. Mulligan & A. E. Porsild) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) |
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