Physaria chambersii |
Physaria bellii |
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Bell's or Front Range twinpod, Bell's twinpod, Front Range twinpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex simple, (relatively large); densely (silvery) pubescent, trichomes (sessile, appressed), rays furcate, fused at base. |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
simple from base, decumbent to nearly prostrate, 0.5–1.3 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
(strongly rosulate; shortly petiolate); blade broadly obovate, 1.5–7.5 (width 7.5–26 mm, base gradually tapering to petiole), margins shallowly dentate, (apex obtuse). |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
blade oblanceolate to broadly obovate, 1–2.5 cm, margins entire. |
Racemes | congested. |
dense. |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (pale yellow or yellow-green), narrowly lanceolate to narrowly deltate, 4–8 mm; petals yellow, broadly spatulate to obovate, 9–13 mm, (not clawed). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(divaricate-ascending to widely spreading, slightly sigmoid to curved), 7–12 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
didymous, slightly flattened (contrary to replum) to uncompressed, 4–9 × 2–8 mm, (strongly coriaceous, apical and basal sinuses narrow, deep); valves (retaining seeds after dehiscence), pubescent, trichomes appressed; replum narrowly oblanceolate to narrowly linear-oblong, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style more than 3 mm. |
Seeds | flattened. |
compressed. |
2n | = 8, 10, 16, 24. |
= 8. |
Physaria chambersii |
Physaria bellii |
|
Phenology | Flowering Apr–Jul. | Flowering Mar–Jun(-Jul). |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Dark shale, road cuts, ridge crests, washes |
Elevation | 1500-3200 m (4900-10500 ft) | 1500-1800 m (4900-5900 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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CO
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria bellii is often found in shale and limestone soils of the Fountain/Ingleside, Lykins, Niobrara, and Pierre formations. It is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 628. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | G. A. Mulligan: Canad. J. Bot. 44: 1662, fig. 1, plate 1, fig. 3. (1966) |
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