Physaria chambersii |
Physaria aurea |
|
---|---|---|
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
golden bladderpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Biennials or perennials; (short-lived); caudex branched; densely pubescent, trichomes (sessile or short-stalked, simple or not), 5–9-rayed, rays furcate, (fine, smooth or finely tuberculate). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
several from base, erect or outer ones decumbent or procumbent, (sometimes much-branched distally), to 6 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade obovate or rhombic, to ca. 2.5 cm, margins usually shallowly dentate, sometimes lyrate-pinnatifid. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(proximal shortly petiolate, distal sessile); blade obovate to rhombic or oblanceolate, 2–4(–6) cm, margins entire or shallowly and remotely dentate. |
Racemes | congested. |
usually dense, (several-flowered). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals ovate or oblong (tapering at base), 3.6–4.8(–5.3) mm, (lateral pair subsaccate, median pair thickened apically, cucullate); petals obovate to spatulate, 4.5–7.5 mm, (blade narrowed to broad claw, margins sinuate). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(strongly recurved), to 20 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(± pendent), ovoid, obcompressed, or globose, compressed, 4–6(–8) mm; valves (not retaining seeds after dehiscence), sparsely pubescent or glabrous, sparsely pubescent inside; replum as wide as or wider than fruit; ovules usually 4, rarely 6, per ovary; style 2.5–3.6 mm. |
Seeds | flattened. |
flattened. |
2n | = 8, 10, 16, 24. |
= 14. |
Physaria chambersii |
Physaria aurea |
|
Phenology | Flowering Apr–Jul. | Flowering Jun–Aug. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Open sites and bare areas in rocky limestone soil in mountains, roadbanks, open woods |
Elevation | 1500-3200 m (4900-10500 ft) | 2000-2800 m (6600-9200 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
|
NM |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria aurea (known from the Jicarilla and Sacramento mountains) is similar to 35. P. gooddingii, which is found farther west in the mountains of Catron and Sierra counties, New Mexico, and Greenlee County, Arizona. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 628. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella aurea |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (Wooton) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) |
Web links |