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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod

Great Plains bladderpod

Habit Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). Perennials or, rarely, annuals; caudex simple or branched; ± densely pubescent, trichomes (sessile or short-stalked), few-rayed, rays (usually spreading), distinct or slightly fused at base, furcate or bifurcate, (tuberculate).
Stems

several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm.

simple or few from base, prostrate or straggling to erect, (sometimes purplish, usually unbranched), (0.5–)1–2(–3) dm.

Basal leaves

(petiole slender);

blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate.

blade oblanceolate, 1.5–5(–7) cm, margins entire or shallowly dentate, (flat).

Cauline leaves

blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute).

blade elliptic to linear, (0.5–)1–2.5(–3) cm, margins usually entire.

Racemes

congested.

(secund), dense, (elongated in fruit).

Flowers

sepals narrowly lanceolate, 5–8(–9) mm;

petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade).

sepals elliptic or oblong, 4–6(–7) mm, (lateral pair subsaccate, median pair thickened apically, cucullate);

petals (often red or lavender when dried), obovate, 6–8.5(–9.5) mm, (narrowing to broad claw).

Fruiting pedicels

(divaricate, slightly sigmoid), 8–15 mm.

(usually sharply recurved, sometimes divaricate-spreading or nearly horizontal), 5–15(–20) mm, (stout).

Fruits

(often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded);

valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent;

replum oblong, as wide as or wider than fruits, apex obtuse;

ovules 4–12 per ovary;

style (4–)6–8 mm (exceeding sinus).

subglobose, obovoid, or broadly ellipsoid, slightly inflated, (3.5–)4–5.5(–6.5) mm;

valves densely pubescent outside, trichomes spreading or closely appressed, rarely sparsely pubescent inside;

ovules (4–)8(–10) per ovary;

style (slender), 3–5.5(–6.5) mm.

Seeds

flattened.

slightly flattened.

2n

= 8, 10, 16, 24.

Physaria chambersii

Physaria arenosa

Phenology Flowering Apr–Jul.
Habitat Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas
Elevation 1500-3200 m (4900-10500 ft)
Distribution
from FNA
AZ; CA; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; MT; ND; NE; SD; WY; AB; MB; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Perennials or, rarely, annuals, short-lived; fruit valves: trichomes spreading.
subsp. arenosa
1. Perennials, long-lived; fruit valves: trichomes closely appressed.
subsp. argillosa
Source FNA vol. 7, p. 631. FNA vol. 7, p. 626.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. arenosa subsp. arenosa, P. arenosa subsp. argillosa
Synonyms P. chambersii var. canaani, P. chambersii var. sobolifera Vesicaria arenosa, Lesquerella arenosa, Lesquerella argentea var. arenosa, Lesquerella ludoviciana var. arenosa
Name authority Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) (Richardson) O’Kane & Al-Shehbaz: Novon 12: 321. (2002)
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