Physaria chambersii |
Physaria angustifolia |
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
threadleaf bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Annuals; with a fine taproot; ± densely pubescent, trichomes several-rayed, rays distinct or fused at base, bifurcate, (prominently tuberculate throughout). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
simple or few to several from base, erect, (sometimes branched), to 4 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade elliptic to rhombic, 3–8 cm, (base narrowing gradually to petiole), margins entire, repand, coarsely toothed, or pinnatifid. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(proximal often shortly petiolate, distal sessile); blade linear or narrowly obovate, 1.5–6(–10) cm, margins entire, repand, or shallowly toothed. |
Racemes | congested. |
usually loose. |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals elliptic or ovate, 4–6 mm, (lateral pair usually subsaccate); petals obovate to obdeltate, 6–10 mm, (apex often emarginate). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(usually divaricate, sometimes horizontal, straight or slightly curved), 8–20 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
not didymous, ± globose, slightly inflated, 4–6 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4 per ovary; style 2–3.5 mm; (stigma expanded). |
Seeds | flattened. |
flattened, (margined). |
2n | = 8, 10, 16, 24. |
= 10. |
Physaria chambersii |
Physaria angustifolia |
|
Phenology | Flowering Apr–Jul. | Flowering Apr(-May). |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Shallow limestone-derived soils, sometimes spreading to disturbed sites |
Elevation | 1500-3200 m (4900-10500 ft) | 90-300 m (300-1000 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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OK; TX |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 625. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Vesicaria angustifolia, Lesquerella angustifolia, Lesquerella longifolia |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (Nuttall ex Torrey & A. Gray) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) |
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