Physaria chambersii
Physaria acutifolia
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod
double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod
several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm.
several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm.
(petiole slender);
blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate.
(petiole slender, often narrowly winged);
blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro).
blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute).
blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse).
congested.
loose, (elongated in fruit).
sepals narrowly lanceolate, 5–8(–9) mm;
petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade).
sepals linear-oblong, 4–7.5 mm;
petals spatulate, 6–11 mm.
(divaricate, slightly sigmoid), 8–15 mm.
(divaricate, slightly sigmoid or nearly straight), 6–12 mm.
(often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded);
valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent;
replum oblong, as wide as or wider than fruits, apex obtuse;
ovules 4–12 per ovary;
style (4–)6–8 mm (exceeding sinus).
(erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so);
valves retaining seeds after dehiscence, pubescent, trichomes appressed;
replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse;
ovules (2 or) 4 per ovary;
style 4–6(–9) mm.
flattened.
(dark brown), flattened, (2–3 mm).
= 8, 10, 16, 24.
= 10, 16, 24.
Physaria chambersii
Physaria acutifolia
Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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