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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod

Habit Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). Annuals, biennials, perennials, or subshrubs; eglandular.
Stems

several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm.

Basal leaves

(petiole slender);

blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate.

Cauline leaves

blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute).

petiolate, sessile, or subsessile;

blade base usually not auriculate (except Paysonia), margins entire, dentate, or sinuate.

Racemes

congested.

ebracteate, often elongated in fruit.

Flowers

sepals narrowly lanceolate, 5–8(–9) mm;

petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade).

actinomorphic;

sepals erect, spreading, ascending, or reflexed, lateral pair seldom saccate basally;

petals white, yellow, lavender, purple, violet, orange, or brown [pink], claw present, often distinct;

filaments unappendaged, not winged;

pollen (3 or) 4–11-colpate.

Fruiting pedicels

(divaricate, slightly sigmoid), 8–15 mm.

Fruits

(often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded);

valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent;

replum oblong, as wide as or wider than fruits, apex obtuse;

ovules 4–12 per ovary;

style (4–)6–8 mm (exceeding sinus).

silicles or siliques, dehiscent, unsegmented, terete, latiseptate, or angustiseptate;

ovules 2–100 per ovary;

style usually distinct;

stigma entire or strongly 2-lobed.

Seeds

flattened.

biseriate, uniseriate, or aseriate;

cotyledons accumbent or incumbent.

Trichomes

usually short-stalked, subsessile, or sessile, sometimes long-stalked, stellate, scalelike, subdendritic, or forked, sometimes mixed with simple ones.

2n

= 8, 10, 16, 24.

Physaria chambersii

Brassicaceae tribe Physarieae

Phenology Flowering Apr–Jul.
Habitat Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas
Elevation 1500-3200 m (4900-10500 ft)
Distribution
from FNA
AZ; CA; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
North America; Mexico; South America; Asia (ne Russia)
Discussion

Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 7, species ca. 130 (7 genera, 105 species in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 631. FNA vol. 7, p. 604.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
Synonyms P. chambersii var. canaani, P. chambersii var. sobolifera
Name authority Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) B. L. Robinson: in A. Gray et al., Syn. Fl. N. Amer. 1(1,1): 100. (1895)
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