Physaria chambersii |
Brassicaceae tribe Physarieae |
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Annuals, biennials, perennials, or subshrubs; eglandular. |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
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Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
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Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
petiolate, sessile, or subsessile; blade base usually not auriculate (except Paysonia), margins entire, dentate, or sinuate. |
Racemes | congested. |
ebracteate, often elongated in fruit. |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
actinomorphic; sepals erect, spreading, ascending, or reflexed, lateral pair seldom saccate basally; petals white, yellow, lavender, purple, violet, orange, or brown [pink], claw present, often distinct; filaments unappendaged, not winged; pollen (3 or) 4–11-colpate. |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
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Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
silicles or siliques, dehiscent, unsegmented, terete, latiseptate, or angustiseptate; ovules 2–100 per ovary; style usually distinct; stigma entire or strongly 2-lobed. |
Seeds | flattened. |
biseriate, uniseriate, or aseriate; cotyledons accumbent or incumbent. |
Trichomes | usually short-stalked, subsessile, or sessile, sometimes long-stalked, stellate, scalelike, subdendritic, or forked, sometimes mixed with simple ones. |
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2n | = 8, 10, 16, 24. |
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Physaria chambersii |
Brassicaceae tribe Physarieae |
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Phenology | Flowering Apr–Jul. | |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | |
Elevation | 1500-3200 m (4900-10500 ft) | |
Distribution |
AZ; CA; ID; NV; OR; UT
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North America; Mexico; South America; Asia (ne Russia) |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 7, species ca. 130 (7 genera, 105 species in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 604. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | B. L. Robinson: in A. Gray et al., Syn. Fl. N. Amer. 1(1,1): 100. (1895) |
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