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Idaho bladderpod

largefruit bladderpod

Habit Perennials; caudex simple, (often enlarged by persistent leaf bases); densely pubescent, trichomes (sessile or short-stalked), rays furcate or bifurcate, (nearly smooth to finely tuberculate). Perennials; caudex branched; densely pubescent, trichomes (sessile or short-stalked), 4–6-rayed, rays distinct, usually furcate, rarely bifurcate, (finely tuberculate throughout).
Stems

simple from base, decumbent, (occasionally few-branched), 0.5–1.5(–2) dm.

few or several from base, prostrate to decumbent, (unbranched or branched), 0.5–1.5 dm.

Basal leaves

blade elliptic to broadly obovate, triangular, rhombic, or round, 1.5–3(–4) cm, margins often sinuate or shallowly lobed.

blades orbicular to broadly obovate, 1.5–3 cm, margins usually entire, rarely remotely dentate.

Cauline leaves

(sessile or shortly petiolate);

blade elliptic to oblanceolate to obovate, 0.5–1.5 cm, (base narrowed to petiole), margins entire.

(sessile or shortly petiolate);

blade elliptic to oblanceolate, 1–1.5(–2.5) cm, margins entire, (apex obtuse).

Racemes

compact to loose.

dense, (elongated in fruit).

Flowers

sepals oblong to broadly elliptic, 4–7.5 mm, (lateral pair saccate or not);

petals spatulate, 7.5–10 mm.

sepals ovate or oblong-elliptic, 5–5.5 mm, (lateral pair not saccate);

petals cuneate or broadly obovate, ca. 7 mm, (sometimes slightly narrowed to a broad claw, apex sometimes retuse).

Fruiting pedicels

(ascending or divaricate-spreading, straight to loosely sigmoid or curved), 4–10 mm.

(sharply recurved), 5–10 mm, (stout).

Fruits

(sessile or substipitate), elliptic, suborbicular, or oblong, strongly compressed (angustiseptate), 5–9 mm, (rounded to sharply keeled on 1 side, edges ± keeled);

valves: (margins covering replum edges or not), usually pubescent throughout or, rarely, glabrous inside;

ovules (4–)8–14(–16) per ovary;

style 2–4.5(–5) mm.

subglobose to broadly obovoid, strongly inflated (often slightly angustiseptate), 5–7 mm, (papery);

valves sparsely pubescent; (septum fenestrate, perforate, or obsolete);

ovules 4–8 per ovary;

style 2–3 mm.

Seeds

slightly flattened.

somewhat flattened.

Physaria carinata

Physaria macrocarpa

Phenology Flowering May–Jun.
Habitat Gypsum-clay hills and benches, naked clay flats and barren hills
Elevation 2000-2400 m (6600-7900 ft)
Distribution
from FNA
ID; MT; WY
[BONAP county map]
from FNA
WY
[BONAP county map]
Discussion

Subspecies 3 (3 in the flora).

Differences in fruit morphology become blurred and the three subspecies are often indistinguishable where their ranges meet near the intersection of Idaho, Montana, and Wyoming.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Physaria macrocarpa is found in the Great Divide and Green River basins.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruits elliptic, not keeled, valve margins (thin, rounded), not covering replum edges.
subsp. paysonii
1. Fruits elliptic, suborbicular, or broadly oblong, keeled on one side, valve margins covering replum edges
→ 2
2. Fruits elliptic, bases narrow-rounded to acute, apices ± acute.
subsp. carinata
2. Fruits suborbicular to elliptic or broadly oblong, bases and apices rounded.
subsp. pulchella
Source FNA vol. 7, p. 629. FNA vol. 7, p. 650.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. carinata subsp. carinata, P. carinata subsp. paysonii, P. carinata subsp. pulchella
Synonyms Lesquerella carinata Lesquerella macrocarpa
Name authority (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) (A. Nelson) O’Kane & Al-Shehbaz: Novon 12: 325. (2002)
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